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Geologic formation of Early Cretaceous age in northeastern Brazil
The Crato Formation is a geologic formation of Early Cretaceous (Aptian) age in northeastern Brazil's Araripe Basin. It is an important Lagerstätte (undisturbed
Crato_Formation
Stratigraphic Group in Brazil
comprises the Crato, Ipubi and Romualdo Formations and is dated to the Aptian to Albian stages of the Early Cretaceous. The formations of the group were
Santana_Group
morphology and taphonomy of elcanid fossils from the Lower Cretaceous Crato Formation (Brazil), interpret the studied insects as living close to the lake
2026_in_paleoentomology
Brazilian geologic formation
ultimately from Romualdo Formation rocks. The Crato Formation was previously considered the lowest member of the then Santana Formation, but has been elevated
Romualdo_Formation
Extinct genus of centipedes
centipede. The Nova Olinda Member of the Early Cretaceous Brazilian Crato Formation, where the remains were found, is known for its excellent preservation
Cratoraricrus
Family of crickets
Martins-Neto 1987 Crato Formation, Brazil, Aptian ?Weald Clay, United Kingdom, Hauterivian †Brontogryllus Martins-Neto 1991 Crato Formation, Brazil, Aptian
Gryllidae
Informal species of theropod
known by a single species, "Ubirajara jubatus", recovered from the Crato Formation. It was described as the first Gondwanan non-avian theropod dinosaur
Ubirajara_jubatus
Genus of Malacostraca
Karl Beurlen (1901–1985). Fossils of the shrimp were found in the Crato Formation of the Araripe Basin of northeastern Brazil. A study on the anatomy
Beurlenia
Genus of tapejarid pterosaur from the Early Cretaceous
genus of tapejarid pterodactyloid pterosaur from the Early Cretaceous Crato Formation of Brazil. It is known from two species, T. imperator and T. navigans
Tupandactylus
Single-species extinct genus of moths
Eolepidopterigidae, containing one species, Psamateia calipsa, which is known from the Crato Formation in Brazil. Insects of the Crato Formation v t e v t e
Psamateia
Family of insects
Martins-Neto & Vulcano, 1988 Crato Formation, Brazil, Early Cretaceous (Aptian) †Baisochrysa Makarkin, 1997 Karabastau Formation, Kazakhstan, Middle/Late
Chrysopidae
Extinct genus of lizard
extinct genus of squamate from the Early Cretaceous (Aptian) aged Crato Formation of Brazil. It has an elongated snake-like body, with four disproportionately
Tetrapodophis
Extinct genus of ray-finned fishes
cylindricus Agassiz, 1841 - Late Aptian/Early Albian of Brazil (Crato and Santana Formations) †C. mawsoni Woodward, 1902 - Late Hauterivian/Early Barremian
Calamopleurus
Members of the insect family Gryllotalpidae
†Archaeogryllotalpoides Martins-Neto 1991 Crato Formation, Brazil, Aptian †Cratotetraspinus Martins-Neto 1997 Crato Formation, Brazil, Aptian †Marchandia Perrichot
Mole_cricket
Order of arachnids known as whip scorpions
reach 85 mm (3.3 in). An extinct Mesoproctus from the Lower Cretaceous Crato Formation could be the same size. Because of their legs, claws, and "whip", though
Uropygi
Topics referred to by the same term
Crato may refer to: Crato, Portugal Crato, Ceará, Brazil Crato, Count of Nassau-Saarbrücken (1621–1642), German nobleman Louis Crato, Count of Nassau-Saarbrücken
Crato
Subclass of plants containing three genera of gymnosperms
Priscowelwitschia Dilcher et al., 2005 Crato Formation, Brazil, Early Cretaceous (Aptian) Cratonia Rydin et al., 2003 Crato Formation, Brazil, Early Cretaceous (Aptian)
Gnetidae
Genus of tapejarid pterosaur from the Early Cretaceous
finger") is a genus of tapejarid pterosaur from the Early Cretaceous Crato Formation of Brazil. It contains a single species, A. cearensis. The holotype
Aymberedactylus
Extinct genus of moths
Parasabatinca is an extinct genus of small primitive metallic moths within the extant family Micropterigidae or extinct family Eolepidopterigidae, containing
Parasabatinca
Extinct genus of coelacanths
nodules. These fossils come from the Crato Formation, Aptian in age, and especially from the Romualdo Formation, whose age is generally considered Albian
Axelrodichthys
Extinct genus of fungi
of gilled fungus in the order Agaricales from the Early Cretaceous Crato Formation of Brazil. It contains the single species G. magnificus, and it is
Gondwanagaricites
Extinct genus of dinosaurs
record of the group at South America. The genus was named after the Crato Formation, in which the specimen was found. The holotype is associated with elements
Cratoavis
Extinct genus of praying mantises
Santanmantidae. The only species, Santanmantis axelrodi , is known from the Crato Formation of Brazil, dating to the late Aptian stage of the Early Cretaceous
Santanmantis
Geologic formation in Brazil
northeastern Brazil. The formation is dated to the Aptian to Albian stages of the Early Cretaceous, unconformably overlying the Crato Formation and unconformably
Ipubi_Formation
Extinct genus of insects
Cretoraphidia, Cretoraphidiopsis and Lugala. Its fossils were found at the Crato MNHN collection in Brazil, often known as the Cretaceous of Brazil. It contains
Austroraphidia
Family of plants
southwestern Africa. Three fossil genera have been recovered from the Crato Formation – late Aptian (Lower Cretaceous) strata located in the Araripe Basin
Welwitschiaceae
Extinct genus of arachnids
Britopygus is an extinct genus of whip spider that lived in Brazil (Crato Formation) during the Early Cretaceous epoch. The genus is known from a single
Britopygus
Extinct family of moths
†Eolepidopterix Rasnitsyn, 1983 Uda Formation, Russia, Oxfordian †Gracilepteryx Martins-Neto and Vulcano, 1989 Crato Formation, Brazil, Aptian †Grammikolepidopteron
Eolepidopterigidae
Extinct genus of fishes
Jordan, 1910 from the Aptian of the Muribeca Formation (Sergipe-Alagoas Basin) and the Crato Formation, where it is an abundant fossil fish. The species
Dastilbe
Genus of fishes (fossil)
England, where they are known from the Toarcian-aged Whitby Mudstone Formation. "Lepidotes" latifrons from the Middle-Late Jurassic of England has been
Lepidotes
Extinct genus of insects
recovered from outcrops of the Nova Olinda Member of the Late Aptian Crato Formation in Northeastern Brazil. P. rudgwickensis was described from a single
Principiala
Extinct genus of spider-like animals
Cratosolpuga wunderlichi. The genus is known from and named after the Crato Formation. Paleontology portal "†Cratosolpuga Selden 1996 (camel spider)". The
Cratosolpuga
Extinct genus of amphibians
Cretaceous (Aptian). Fossils were discovered in and named after the Crato Formation. It was named by Ana M. Báez, Geraldo J.B. Moura and Raúl O. Gómez
Cratia
Extinct genus of lizards
America. The type species Calanguban alamoi was named in 2014 from the Crato Formation of Brazil and is the oldest known non-iguanian lizard from the continent
Calanguban
Extinct genus of Brazilian ant
species, Vulcanidris cratensis. The genus was first discovered in the Crato Formation of Northeastern Brazil, and it represents the oldest verified ant known
Vulcanidris
Aquatic insects of the order Ephemeroptera
Cretereismatidae from the Lower Cretaceous Crato Formation of Brazil also belongs as the last offshoot to Permoplectoptera. The Crato outcrops otherwise yielded fossil
Mayfly
Extinct plant genus
Cretaceous Crato Formation of Brazil. A probable occurrence of this genus has also been reported from the Late Cretaceous Akrabou Formation of southeast
Welwitschiophyllum
Genus of anhanguerid pterosaur from the Early Cretaceous
from the Early Cretaceous period (Aptian stage) of what is now the Crato Formation of the Araripe Basin in Ceará, Brazil. The type and only species is
Ludodactylus
Extinct genus of ray-finned fishes
was found in the Crato and Santana Formations. Other fossils of the genus have been found at Goulmima in Morocco, the Tlayua Formation of Mexico and the
Araripichthys
Extinct genus of ray-finned fishes
and Crato Formations of the Araripe Basin) (=C. ferus Santos, 1950) †C. geddesi Berell, 2014 - late Albian of Queensland, Australia (Winton Formation) An
Cladocyclus
Extinct family of grasshoppers
Cratolocustopsis Martins-Neto, 2003 Crato Formation, Brazil, Aptian † Cratozeunerella Martins-Neto, 1998 Crato Formation, Brazil, Aptian † Liadolocusta Handlirsch
Locustopsidae
Single-species extinct genus of moths
is known from the Late Aptian Crato Formation of the Araripe Basin in northeastern Brazil. "Insects of the Crato Formation" (PDF). Archived from the original
Gracilepteryx
Geological formation in Australia
Group, contemporaneous Lagerstätte in northeastern Brazil Crato Formation Romualdo Formation Weishampel et al., 2004, pp.573-574 Frauenfelder TG, Bell
Toolebuc_Formation
Genus of thalassodromid pterosaur from the Early Cretaceous
million years ago. Its remains were found in what is now the Romualdo Formation of the Santana Group in Brazil. Tupuxuara was named in 1988 by paleontologists
Tupuxuara
Suborder of insects
Timematodea. The oldest record of Euphasmatodea is Araripephasma from the Crato Formation of Brazil, dating to the Aptian stage of the Early Cretaceous. The
Euphasmatodea
Extinct family of true bugs
palaeontinid fossils include the Crato Formation Lagerstätte of Brazil and the Yixian Formation, Haifanggou (or Jiulongshan) Formation, and the Daohugou Beds of
Palaeontinidae
the subfamily Anophelinae, is described from the Lower Cretaceous Crato Formation (Brazil) by Oliveira et al. (2025). Redescription of Aetheapnomyia
2025_in_paleoentomology
Genus of semionotiform fish
the Araripe Basin, and was commonly found in Santana formation, and rare in the Crato Formation, mostly in carbonate concretions, but uncommon in laminated
Araripelepidotes
Extinct genus of insects
of South America and Asia. The type species is A. seldeni from the Crato Formation of Brazil (Araripe Basin). The two other species are A. burmanus from
Araripenymphes
Genus of tapejarid pterosaur
M. (2007). "Pterosaurs of the Crato Formation." In Martill, D. M., Bechly, G. and Loveridge, R. F. (eds), The Crato Fossil Beds of Brazil: Window into
Tapejara_wellnhoferi
Family of mayflies
al. 2020 Crato Formation, Brazil, Early Cretaceous (Aptian) Subfamily †Colocrurinae McCafferty, 1990 †Colocrus McCafferty 1990 Crato Formation, Brazil
Oligoneuriidae
Family of true bugs
"water scorpions". †Araripenepa from the Early Cretaceous (Aptian) Crato Formation of Brazil is the oldest known member of the family, and is the sister
Nepidae
Family of dragonflies
Cenomanian †Cratohagenius Bechly, 2010 Crato Formation, Brazil, Aptian †Cratolindenia Bechly, 2000 Crato Formation, Brazil, Aptian †Gunterbechlya Huang
Gomphidae
Order of stick and leaf insects
oldest known being Araripephasma from the Early Cretaceous (Aptian) Crato Formation of Brazil, around 113 million years old, which can be confidently assigned
Phasmatodea
Family of plants
Arlenea Ribeiro, Yang, Saraiva, Bantim, Calixto Junior et Lima, 2023 Crato Formation, Brazil, Early Cretaceous (Aptian) Leongathia V.A. Krassilov, D.L.
Ephedraceae
Extinct genus of dragonflies
the Early Cretaceous. It contains nine species; most are from the Crato Formation of Brazil, though C. europaeus is from the Weald Clay of England. C
Cordulagomphus
Extinct family of dragonflies
aeschnidiid dragonfly genus and species from the Lower Cretaceous Crato Formation (Odonata, Anisoptera). Historical Biology 35:865-869 A. Nel. 2021.
Aeschnidiidae
Family of spiders
Cretaceous (Aptian) Crato Formation, Brazil †Cretadiplura ceara Selden, 2005 †Dinodiplura Selden, 2005 — Early Cretaceous (Aptian) Crato Formation, Brazil †Dinodiplura
Dipluridae
Genus of azhdarchoid pterosaur from the Early Cretaceous
stage, about 120 to 93 million years ago) Nova Olinda Member of the Crato Formation. The skull was long, and unusually wide. The section in front of the
Lacusovagus
Order of arachnids
the Triassic. So far, they are also absent from the Lower Cretaceous Crato Formation of Brazil, a Lagerstätte that has yielded many other terrestrial arachnids
Opiliones
Geological formation in Australia
Group, contemporaneous Lagerstätte in northeastern Brazil Crato Formation Romualdo Formation South Polar region of the Cretaceous Weishampel et al., 2004
Allaru_Formation
Order of insects
Willmann, 1994 - Crato Formation, Brazil, Early Cretaceous (Aptian) Genus †Baissoptera Martynova, 1961 - Crato Formation, Brazil, Yixian Formation, China, Zaza
Snakefly
Family of centipedes
†Fulmenocursor, an extinct genus from the Early Cretaceous of the Crato Formation of Brazil. Rosario Dioguardi. "Chilopoda taxonomic browser". ChiloBase
Scutigeridae
Extinct genus of insects
cratoensis specimen was collected from layers of the Upper Aptian Crato Formation. The formation is composed of unweathered grey and oxidized yellow limestones
Rafaelnymphes
Genus of ornithocheirid pterosaur from the Early Cretaceous
Arthurdactylus is a genus of pterodactyloid pterosaur from the Early Cretaceous Crato Formation of northeastern Brazil. It was a medium-sized pterosaur, with a wingspan
Arthurdactylus
Family of insects
Martins-Neto 1992 – Crato Formation, Brazil, Aptian Derhynchia Halter Halterina Knersvlaktia †Krika Martins-Neto 1992 – Crato Formation, Brazil, Aptian Lertha
Nemopteridae
Single-species extinct genus of moths
Eolepidopterigidae, containing one species, Netoxena nana, which is known from the Crato Formation of the Araripe Basin in northeastern Brazil. Its generic name was originally
Netoxena
Extinct genus of insects
Kalligrammatidae described by Martins-Neto in 1997 from fossils found in the Crato Formation of the Araripe Basin in northeastern Brazil. The genus contains three
Makarkinia
Superfamily of insects
amber, Cenomanian †Lalacidae Hamilton 1990 Crato Formation, Brazil Lushangfen Formation, Yixian Formation, China, Aptian †Mimarachnidae Shcherbakov 2007
Planthopper
Limestone containing fossils
Humankind Crato Formation Cumnor Hurst Cunswick Scar Dinosaur Valley State Park Durupınar site Dudley Tunnel East Kirkton Quarry Elliot Formation English
Fossiliferous_limestone
Clade of amphibians
Candeleros Formation, Late Cretaceous (Cenomanian) †Vulcanobatrachus Trueb et al. 2005 South Africa, Late Cretaceous †Cratopipa Carvalho et al., 2019 Crato Formation
Pipoidea
Genus of evergreen conifers
jeffreyi – Eastern United States (Late Cretaceous) †Araucaria benignae – Crato Formation, Brazil (Early Cretaceous (Aptian)) Araucaria bindrabunensis (previously
Araucaria
Extinct genus of moths
Cretaceous. The second species, Undopterix cariensis is known from the Crato Formation in Brazil. Undopterix was initially classified in the Micropterigidae
Undopterix
Extinct family of insects
myrmeleontodoids, such as antlions. †Babinskaia Martins-Neto and Vulcano 1989 Crato Formation, Brazil, Aptian †Babinskaia formosa Martins-Neto and Vulcano 1989 †Babinskaia
Babinskaiidae
Extinct genus of turtles
from 112 to 109 million years ago, in the Early Cretaceous Crato and Romualdo Formations of the Araripe Basin in northeastern Brazil. It is one of the
Araripemys
Extinct genus of amphibians
alcinae. Eurycephalella was discovered within the limestone predominant Crato Formation of the Araripe Basin in northeastern Brazil. The specimen is the partial
Eurycephalella
Geological formation in Niger, West Africa
over millions of years. List of dinosaur-bearing rock formations Crato Formation Romualdo Formation SCIENCE IN THE NEWS - Nov. 13: Digest - 12 November
Elrhaz_Formation
Extinct genus of reptiles
Fossils have been found from the Nova Olinda Member of the Aptian-age Crato Formation in the Araripe and Lima Campos Basins of northeastern Brazil. Named
Susisuchus
Extinct clade of lizards
Cryptobicuspidon pachysymphysealis from the Quiricó Formation of Brazil and Calanguban alamoi from the Crato Formation of Brazil, while Early Cretaceous South American
Polyglyphanodontia
Extinct family of insects
Crato Formation, Brazil An. myrmicopterus Rasnitsyn, 1975 - Aptian Baissa, Zaza Formation, Russia An. niger Rasnitsyn, 1990 - Aptian, Turga Formation
Angarosphecidae
Fictional character
Retrieved July 1, 2022. "A New Species of Brachyphyllum from the Crato Formation" (PDF). Ameghiniana. June 28, 2020. Retrieved June 29, 2022. Etymology
Ellie_Sattler
Extinct family of insects
†Dryellina Martins-Neto and Rodrigues 2009 Crato Formation, Brazil, Aptian †Karenina Martins-Neto 1997 Crato Formation, Brazil, Aptian †Kareninoides Yang et
Mesochrysopidae
Order of insects including praying mantises
compression fossils (in rock) include adults. Fossil mantises from the Crato Formation in Brazil include the 10 mm (0.39 in) long Santanmantis axelrodi, described
Mantis
Extinct order of insects
described from Mesozoic outcrops, mainly from the Lower Cretaceous Crato Formation of Brazil (in total, around 40 fossil nymphs have been found). Both
Coxoplectoptera
Spinosaurid theropod dinosaur genus from the Early Cretaceous Period
earlier, Frey and Martill had named a new pterosaur species from the Crato Formation Arthurdactylus conandoylei, after the novelist himself. When Martill
Irritator
Extinct family of cockroaches
1999 Crato Formation, Brazil, Early Cretaceous (Aptian) †Blattapterix Vršanský 2003 Xiagou Formation, China, Early Cretaceous (Aptian); Jinju Formation, South
Umenocoleidae
Rift Basin in brazil famous for its pterosaur fossils
Basins. The basin has provided a variety of unique fossils in the Crato and Santana Formations and includes the Araripe Geopark, a member of the UNESCO Global
Araripe_Basin
Genus of anhanguerian pterosaur
Janeiro Frey, E. & Martill, D. M. (1994). "A new Pterosaur from the Crato Formation (Lower Cretaceous, Aptian) of Brazil." Neues Jahrbuch fur Geologie
Brasileodactylus
Fifth age of the Early Cretaceous
(Cheirolepidiaceae, Coniferales) in the peri-equatorial late Aptian Crato Formation (Santana group, Araripe Basin, Brazil) and their paleoclimatic and
Aptian
Genus of theropod dinosaur
(Schizaeales, Anemiaceae) – A common fern from the Lower Cretaceous Crato Formation of northeast Brazil". Cretaceous Research. 54: 17–26. Bibcode:2015CrRes
Deinonychus
Organelle found in animal cells used for the synthesis, storage and transport of melanin
from Tupandactylus cf. imperator pterosaurs in the Lower Cretaceous Crato Formation, in the Araripe Basin, in Brazil. Melanosomes are believed to template
Melanosome
Group of insects
anatomical features first appeared in the Lower Cretaceous of the Crato Formation in Brazil, some 65 million years before the first fig trees. The Vespidae
Wasp
Order of arachnids
Kraepelin, 1899 (1 species) †Britopygus Dunlop & Martill, 2002 (1 species; Crato Formation, Brazil, Cretaceous) Heterophrynus Pocock, 1894 (14 species) Paraphrynus
Amblypygi
Family of flies
Kraemer 2014 Mexican amber, Miocene †Megapsychoda Azar and Nel 2002 Crato Formation, Brazil, Early Cretaceous (Aptian) Mormia Enderlein, 1935 Neoarisemus
Psychodidae
Paleontological site in New South Wales, Australia
this Lagerstätte has been compared to the quality of that found at Crato Formation, Lake Eckfeld, and Libros. Other fossil sites in Australia, such as
McGraths_Flat
Extinct genus of conifers
Caturrita Formation The Caballos Formation of Tolima, Colombia The El Plan Formation of the Department of Francisco Morazan, Honduras The Crato Formation of
Brachyphyllum
Family of insects
†Olindanymphes Martins-Neto, 2005 - Crato Formation, Brazil, Early Cretaceous (Aptian) †Santananymphes Martins-Neto, 2005 - Crato Formation, Brazil, Early Cretaceous
Nymphidae
Subfamily of ants
genera) †Myrmeciites Archibald, Cover & Moreau, 2006 The Late Aptian Crato Formation genus Cariridris was sometimes included in Myrmeciinae, but it has
Myrmeciinae
Species of aquatic plant
Oliveira and David Winship Taylor South of Nova Olinda, Brazil in the Crato Formation of the Araripe Basin. It is stored in the Natural History Museum, Berlin
Pluricarpellatia
Fossil Collection
Cretaceous Crato Formation Tropeognathus mesembrinus skeleton 110 Ma Early Cretaceous Santana Formation Dastilbe crandalli 105-110 Ma Early Cretaceous Crato &
Collection of fossils in the National Museum of Brazil
Collection_of_fossils_in_the_National_Museum_of_Brazil
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CRATO FORMATION
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CRATO FORMATION
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