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Example of an organism used to describe its species
A holotype is the single physical example (or illustration) of an organism that was used when the species (or lower-ranked taxon) was formally described
Holotype
Spinosaurid theropod dinosaur genus from the Early Cretaceous Period
announcement of return to Brazil in 2026. In 1996, the specimen became the holotype of the type species Irritator challengeri. The genus name comes from the
Irritator
Extinct genus of dinosaurs
Oviraptor is only known from a single partial skeleton regarded as the holotype, as well as a nest of about fifteen eggs and several small fragments from
Oviraptor
Genus of theropod dinosaur from the Late Jurassic period
Marsh, but more likely was used solely for display. The first specimen, holotype USNM 4735, was discovered and excavated by farmer Marshall Parker Felch
Ceratosaurus
Extinct genus of ungulate
fossa suggest that Andrewsarchus likely had a fairly weak bite force. The holotype of Andrewsarchus mongoliensis is a mostly complete cranium (specimen number
Andrewsarchus
Specimen(s) chosen to represent a newly described taxon
nomenclatural type when there was no holotype specified or the holotype has been lost or destroyed isotype – a duplicate of the holotype syntype – any specimen (or
Type_(biology)
Species of marsupial
Territory. It was identified through genetic studies conducted on the two holotype specimens in 2017. It is only known from 3 museum specimens. It is medium-sized
Arnhem_Plateau_planigale
Taxonomy of a dinosaur genus
coined the name Allosaurus fragilis in 1877 for a fragmentary specimen (the holotype specimen; the specimen on which the species was based). According to Marsh
Taxonomy_of_Allosaurus
Extinct genus of birds
Aguirre-Zabala in Comallo, in the region of Patagonia, and was made the holotype of the new genus and species Kelenken guillermoi in 2007. The genus name
Kelenken
Genus of theropod dinosaurs
Christopher T. Griffin and colleagues published later that year argued that the holotype skull of N. lancensis belonged to a mature individual based on histology
Nanotyrannus
Genus of sauropodomorph dinosaurs
represents the fifth dinosaur genus to be named from Zimbabwe. The Musango holotype specimen consists of five dorsal (trunk) vertebrae, at least two sacral
Musango
Extinct genus of sperm whale from the Miocene epoch
may have had a global presence. The body length of Livyatan melvillei holotype is approximately 13.5–17.5 meters (44–57 ft) long. Comparable to that of
Livyatan
Species of shrub endemic to Australia
area in New South Wales, Australia. It is currently only known from the holotype specimen. Grevillea milleriana is a low, spreading, sub-prostrate shrub
Grevillea_milleriana
Genus of pteranodontoid pterosaur from the Early Cretaceous
Yixian Formation of Liaoning, China. It is known from a single specimen, holotype JZMP-V-12, housed at the Benxi Geological Museum. This specimen was modified
Yixianopterus
Carcharodontosaurid dinosaur genus from the Cretaceous period
Early Cretaceous period, approximately 111 to 106 million years ago. The holotype specimen was discovered in the Candeleros Formation of Patagonia in 1993
Giganotosaurus
Genus of azhdarchid pterosaur
two specimens, the holotype, consisting of a partial postcranial skeleton, and the paratype, consisting of a left humerus. The holotype includes material
Thanatosdrakon
Extinct subspecies of carnivoran
lower carnassial (m1) tooth excavated from deposits in Kuruwita as the holotype. It was recorded as specimen BMNH M51883 in the British Museum Natural
Panthera_leo_sinhaleyus
subrectangular in outline. The holotype was once tentatively assigned to "Coelosaurus" antiquus. Shark bites present on the holotype suggest that the specimen's
List of informally named dinosaurs
List_of_informally_named_dinosaurs
Specimen that helps define what a taxon represents
name of a species and other taxon actually represents, but it is not the holotype (and in botany is also neither an isotype nor a syntype). Often there is
Paratype
Extinct genus of ants
slightly smaller in size then the coeval species. The Titanomyrma lubei holotype was discovered by Louis Lube at DMNS locality 784, the Farson Fish Beds
Titanomyrma
Genus of theropod dinosaur
the holotype specimen. TMP 1994.012.0602 and TMP 1995.005.0001 were referred to this species based on shared anatomical features with the holotype. Two
Tromerosaurus
Genus of theropod dinosaurs
in the Nemegt Formation of Mongolia. In 1970, this specimen became the holotype of the only species within the genus, Deinocheirus mirificus; the genus
Deinocheirus
Genus of reptiles (fossil)
Canada. Gryphoceratops is known only from the holotype ROM 56635, a partial right dentary. The holotype was collected in the northwest corner of Dinosaur
Gryphoceratops
Species of gastropod
whorls, becoming slightly wavy and less pronounced on later ones. The holotype of the species (MNHN IM-2013-19792), measured 27.15 mm. This species occurs
Tosapusia_turriformis
Genus of theropod dinosaurs
Surrey, England, in sediments of the Weald Clay Formation, and became the holotype specimen of Baryonyx walkeri, named by palaeontologists Alan J. Charig
Baryonyx
Extinct genus of marine reptiles
specimen. Based on several distinctive features, he decided to make it the holotype of a subgenus accompanying a new species of Plesiosaurus, which he named
Pliosaurus
Extinct genus of dinosaurs
indicates a generalist diet with a potentially occasional carnivory. The holotype—and only known specimen—has been determined to represent a young adult
Gigantoraptor
Genus of extinct mammals
femur). The holotype was found in a silt and mudstone bed over a 1.8 m2 (19 sq ft) area. Further excavation recovered most of the holotype's skeleton—most
Ambulocetus
Extinct genus of theropod dinosaurs
least two unrelated taxa. Some of the Eocarcharia material, including the holotype (name-bearing) specimen, likely belongs to a baryonychine spinosaurid.
Eocarcharia
Extinct genus of therapsids
related to mammals than to dinosaurs, the latter being sauropsids. The holotype and only known specimen of Dinosaurus was discovered in the early 1840s
Dinosaurus
Extinct genus of dragonfly-like insects
a body length from head to tail of almost 430 millimetres (17 in). The holotype is held in the Museum of Comparative Zoology. Meganeuropsis americana,
Meganeuropsis
Species of starfish
in reference to the upper surface of the body wall, which is soft. The holotype measures 5.4 centimeters from the center to the end of the arms and 2 centimeters
Mediaster_mollis
Species of beetle
Agra dable is a species of carabid beetle. The holotype was collected in Costa Rica and first described to science in 2002. Agra dable measure 14.5–18
Agra_dable
Extinct genus of reptiles
carolinensis, meaning "Carolina butcher". Two specimens are known, the holotype skull and skeleton NCSM 21558, and the referred humerus NCSM 21623. The
Carnufex
Extinct genus of dinosaurs
University of Kansas, Yale University, and the University of New Orleans. The holotype fossil is less than 1 m (3.3 ft) long, although this specimen appears to
Bambiraptor
Extinct genus of jawless fishes
earliest known members of the vertebrate lineage in the fossil record. The holotype of the type species, Myllokunmingia fengjiaoa — and the only uncontroversial
Myllokunmingia
Extinct chimaeric genus of dinosaurs
D. steini were discovered in a multi-species bonebed. Elements of the holotype and referred specimens were later found to belong to trionychid turtles
Dakotaraptor
Genus of theropod dinosaur
lower levels of the Romualdo Formation. The Aratasaurus museunacionali holotype, MPSC R 2089, was discovered in 2008 in Mina Pedra Branca, a plaster mine
Aratasaurus
Genus of theropod dinosaurs
the longest avialan known. The type specimen of Austroraptor cabazai, holotype MML-195, was recovered in the Bajo de Santa Rosa locality of the Allen
Austroraptor
Genus of ctenochasmatid pterosaur from the Early Cretaceous
The specific name refers to the provenance from the Jiangangshan. The holotype, IG-CAGS-08-07, includes the almost complete skull, the lower jaws, the
Gladocephaloideus
Extinct genus of birds
deposits in Inner Mongolia, while T. peregrinus was found in France. The holotype and only known specimen of Telecrex grangeri, AMNH 2942, is a partial right
Telecrex
Extinct genus of dinosaurs
were. Sinornithosaurus was among the smallest dromaeosaurids, with the holotype measuring 1.2 metres (3.9 ft) long and weighing 3–5 kilograms (6.6–11.0 lb)
Sinornithosaurus
Species of damselfly
refers to the fact that the species existed in the Cretaceous period. The holotype, NHMUK CH 879vii, a fossilized wing, was found n lagoonal sediments in
Steleopteron_cretacicus
Extinct genus of dinosaurs
The genus name honours Gideon Mantell, the discoverer of Iguanodon. The holotype fossil, NHMUK PV R5764, was originally discovered by Reginald Walter Hooley
Mantellisaurus
Genus of Cretaceous cephalopods
traditionally recognized. In 2019, due to the poor preservation of the holotype specimen, Tusoteuthis was theorized to likely be a nomen dubium, and it
Tusoteuthis
Extinct genus of dinosaurs
where it was brought for preparation. In 2004, the specimen became the holotype of the new genus and species Atrociraptor marshalli; the generic name is
Atrociraptor
tenuis, Davis 1887 Type Locality: Sendenhorst (Westphalia, Germany) The Holotype is specimen "GIM-8434". As reported in Dietze's study : "Complete specimen
Telepholis
Species of moth
species was first described by John Stewart Dugdale in 1990. The male holotype specimen, collected on the Glory Track at Bluff Hill, is held in the New
Ctenopseustis_filicis
Extinct genus of plesiosauroids
known from a well-preserved, nearly complete skeleton. The Plesionectes holotype specimen, SMNS 51945, was discovered in 1978 by Gotthilf Fischer in outcrops
Plesionectes
Species of gastropod
gastropod mollusk, in the family Costellariidae, the ribbed miters. The holotype of the species (MNHN IM-2013-19695), measured 35.1 mm. Type Locality :
Tosapusia_bismarckeana
Extinct genus of dinosaur
the fossils and enhanced virtual and long-distance collaboration. The holotype specimen, MPM-PV 1156, consists of a partial skeleton, somewhat preserved
Dreadnoughtus
Species of crustacean
Godzillius robustus is a species of remipede in the genus Godzillius. The holotype and paratype were discovered at the Turks And Caicos Islands. At up to
Godzillius_robustus
Extinct genus of stegosaurian dinosaurs
stegosaurs occurring in the Middle Jurassic of Africa. The Thyreosaurus holotype specimen, HIIUC-BN00, was discovered in 2021 in sediments of the El Mers
Thyreosaurus
Extinct genus of dinosaurs
and described in 2018 by Andrew T. McDonald and Douglas G. Wolfe. The holotype specimen, WSC 16505, consists of fragments of a dorsal rib, complete or
Invictarx
Species of polychaete worm
The species lives in the Sea of Japan off Sado Island, Japan, where the holotype was found living within the internal canals of a sponge of the genus Petrosia
Ramisyllis_kingghidorahi
Genus of carcharodontosaurid theropod dinosaur from the Late Jurassic period
Region of southeastern Tanzania. Veterupristisaurus is known from the holotype specimen MB R 1938, an isolated middle caudal vertebra. Two partially fused
Veterupristisaurus
Extinct genus of mammal
colgatei as the type species, while also erecting the species D. berkeyi. The holotype of D. colgatei is a skull (AMNH 19124), and two lower jaws (AMNH 19003
Didymoconus
Extinct genus of dinosaurs
preservation of the C. canadensis holotype and lack of the parietal bar in C. kaiseni. Following the original assignment of the holotype and other skulls to Mojoceratops
Chasmosaurus
Species of moth
Per Olof Christopher Aurivillius in 1910. It is found in Tanzania. The holotype of this species is illustrated on the web-pages of the Swedish Museum of
Parotis_fasciculata
Species of gastropod
terrestrial pulmonate gastropod mollusk in the family Polygyridae. The holotype was collected on May 24, 1975 from talus accumulation of igneous rock on
Ashmunella_todseni
Ankylosaurid dinosaur genus from the Late Cretaceous Period
teeth of two skulls assigned to A. magniventris differed from those of the holotype specimen in some details, and though he expressed a "considerate temptation"
Ankylosaurus
Genus of theropod dinosaur from the Early Jurassic
preserved were collected in 1942. The most complete specimen became the holotype of a new species in the genus Megalosaurus, named M. wetherilli by Samuel
Dilophosaurus
Extinct species of shrew
on the paleontology of mammals in the southwestern United States. The holotype specimen is deposited at the University of Texas at El Paso Biodiversity
Notiosorex_harrisi
Extinct species of bird
published in 2001. The holotype is in the collection of the Museum of New Zealand Te Papa Tongarewa. It is known from the holotype tarsometatarsus and paratypes
Viti_Levu_giant_pigeon
Extinct genus of dinosaurs
most closely related to Alioramus, the only other known alioramin. The holotype specimen, GM F10004, was unearthed in southern China, Ganzhou, at the Nanxiong
Qianzhousaurus
Extinct mammal genus
P0730 as the holotype specimen. The generic name, Pampaluctor, combines a reference to the Argentina Pampas region in which the holotype was found, with
Pampaluctor
Genus of extinct gansuid birds
birds known to demonstrate semi-aquatic adaptations. The Shuilingornis holotype specimen, LY2022JZ3002, was discovered in sediments of the Jiufotang Formation
Shuilingornis
Extinct genus of reptiles
exhumation. In 1903, Samuel Wendell Williston designated the fossil as the holotype of a new plesiosaur genus and species under the name Brachauchenius lucasi
Brachauchenius
Extinct genus of reptiles
morphology of the holotype specimen's jaws and unique dentition. However, two fossil skulls discovered in 2016 indicate that the holotype skull was badly
Atopodentatus
Place where a name-bearing type specimen was collected
define "type locality" as a regulated term, the collecting locality of the holotype or other original material is routinely referred to by botanists as the
Type_locality_(biology)
Genus of anhanguerian pterosaurs
represents the most complete pterosaur known from Australia. The Haliskia holotype specimen, KK F1426, was discovered by Kevin Petersen in November 2021 within
Haliskia
Genus of crayfishes
Palaeoechinastacus is known from one partial body fossil, which is the holotype, and has been deposited in the National Museum of Victoria in Melbourne
Palaeoechinastacus
Extinct species of reptile
upper arm. The study was illustrated by high quality lithographs. The holotype, NHMUK PV OR 46013, was found in a layer of the Kimmeridge Clay Formation
Dacentrurus
Species of king crab
are typically right-handed. It is named after the Seram Sea where its holotype was found. Lithodes ahyongi, also found in the Seram Sea Takeda, Masatsune;
Lithodes_ceramensis
Extinct genus of dinosaurs
Titanosaurus have also been collected from the Kallakurichi Formation. The holotype vertebrae of Titanosaurus indicus were discovered during an exploration
Titanosaurus
Extinct genus of mammals
Auroracetus, from Aurora (the location the holotype was discovered, Aurora N.C.) and the Latin cetus (whale), is a genus of Pontoporiidae, the river dolphins
Auroracetus
Extinct species of reptile
earliest ankylosaurs known from reasonably complete fossil remains. The holotype was discovered in 1995 at the Bone Cabin Quarry West locality, in Albany
Gargoyleosaurus
Genus of hadrosaurid dinosaurs
Arenysaurini found from this locality, after Ajnabia and Minqaria. The Taleta holotype specimen, MHNM.KH.1557, was discovered in a layer of the Oulad Abdoun Basin
Taleta
Extinct species of reptile
be clenched between the teeth of BMNH R1157 (its holotype) may in fact belong to a victim. The holotype, obtained from the Lower Lias of Lyme Regis, was
Temnodontosaurus_eurycephalus
Extinct genus of theropod dinosaurs
part of the same slab as the holotype; and ELDM V1001, a juvenile estimated to have been eight years younger than the holotype. The fossils are part of the
Yutyrannus
Extinct species of reptile
and dinosaurs. Kadimakara is only known from parts of the skull. The holotype specimen, QMF 6710, includes the rear part of the skull and a fragment
Kadimakara_australiensis
Extinct genus of snakeflies
eighteen fossils, the holotype, three paratype females, ten male paratypes, and three paratypes of indeterminate sex. The holotype, number "USNM 31487"
Archiinocellia
Extinct genus of eurypterid
and Latin arachne meaning "spider", translates to "great spider". The holotype (now stored at the Museum of Paleontology at the National University of
Megarachne
Extinct genus of reptiles
within Atoposauridae, most likely Alligatorellus. The Teylers Museum has a holotype specimen Atoposaurus oberndorfi that was bought by curator J.G.S. van Breda
Atoposaurus
Extinct genus of reptiles
similarities to the holotype of Qiupalong. The foot claws of the new specimens in particular were nearly identical to those of the holotype. As a result, they
Qiupalong
Sauropod dinosaur genus from Late Jurassic-Early Cretaceous Period
mounting, and description of the skeleton were completed in 1965. The holotype specimen (CCG V 20401) consisted of an almost complete and articulated
Mamenchisaurus
Genus of moths
Stegothyris fasciculalis, which is found in Madagascar and South Africa. The holotype of 'Stenia fasciculalis' Zeller, 1852 is illustrated on the web pages of
Stegothyris
Extinct genus of dinosaurs
specific name refers to the Tiantai, where the holotype was discovered. Yueosaurus is known only from the holotype ZMNH M8620, an articulated, partial but well
Yueosaurus
Extinct genus of tetrapods
Carboniferous of West Lothian, Scotland. Kirktonecta is known from the holotype specimen UMZC 2002, a and b, an almost complete skeleton including the
Kirktonecta
Genus of monofenestratan pterosaurs
oldest known monofenestratan pterosaur in the fossil record. The Melkamter holotype specimen, MPEF-PV 11530, was discovered in sediments of the Cañadón Asfalto
Melkamter
Species of extinct macaw
were previously known from St. Croix. Wetmore made the tibiotarsus the holotype specimen of a new macaw species, which he placed in the genus Ara, as Ara
St._Croix_macaw
Genus of ornithopod dinosaur
exposes outcrops of the Weald Clay Formation, is known for yielding the holotype specimen of the spinosaurid theropod Baryonyx walkeri in 1983. During the
Paludohadros
Genus of theropod dinosaurs
from a partial skull and several consecutive vertebrae. The Yuanmouraptor holotype specimen, LFGT-ZLJ0115, was discovered in March 2006 in outcrops of the
Yuanmouraptor
Species of gastropod
on the island of Raiatea in the Society Islands, French Polynesia. The holotype and two other specimens, the only ones recorded so far, were collected
Partula_meyeri
Extinct genus of theropod dinosaurs
overlapping in material with the holotype, was referred to Taurovenator. This was done for several reasons: the holotype fits perfectly with MPCA-Pv 803
Taurovenator
Species of moth
at Little River in Canterbury. It is named after the collector of the holotype, Stuart Lindsay. Lindsay collected two males of the species at Little River
Epichorista_lindsayi
Extinct genus of dinosaurs
it was a herrerasaurid or herrerasaurian. A 2019 redescription of its holotype considered Chindesaurus to be a theropod closely related to Tawa, a slightly
Chindesaurus
Extinct genus of dinosaurs
Provence at the foot of which the animal's remains were unearthed. The holotype specimen, MNHN BO001, consists of the second toe claw of the left foot
Pyroraptor
Extinct population of the brown bear
Southwestern United States and Mexico. The specimen later designated the holotype of U. a. nelsoni was shot by H. A. Cluff at Colonia Garcia, Chihuahua,
Mexican_grizzly_bear
Extinct genus of spiders
Mongolarachnidae in 2015. Mongolarachne jurassica is known only from two fossils, the holotype, specimen number "CNU-ARA-NN2010008" which is a mostly complete adult female
Mongolarachne
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