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Coat pattern in horses
The leopard complex is a group of genetically related coat patterns in horses. These patterns range from progressive increases in interspersed white hair
Leopard_complex
American horse breed noted for spotted color pattern
blindness; the latter has been linked to the leopard complex. Artwork depicting prehistoric horses with leopard spotting exists in prehistoric cave paintings
Appaloosa
Horse coat color pattern
sabinos originates in a white patch, and the roaning is uneven. The leopard complex colors, characteristic of the Appaloosa breed, have several manifestations
Roan_(horse)
Horse coat colors and markings
There are a group of coat patterns caused by the leopard gene complex. Not every horse with leopard genetics will exhibit hair coat spotting. However
Equine_coat_color
Horse coat color
pattern. The leopard complex, related to the Leopard (LP) gene, characterizes the Appaloosa and Knabstrupper breeds with their spotted coats. Leopard is genetically
White_horse
Spotted color pattern
leopard, leopard gecko and the leopard tortoise among some others. The Lp (leopard complex) gene is responsible for the leopard color pattern in horses, which
Leopard_pattern
Species of cat native to Africa and Asia
The leopard (Panthera pardus) is one of the five extant cat species in the genus Panthera. It has a pale yellowish to dark golden fur with dark spots
Leopard
Genetic color in horses
not extend past the throatlatch. Spotting and characteristics of the Leopard complex and the various pinto patterns are not accepted, and body spots of
Palomino
Coloration of animal coat/fur
color variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Points_(coat_color)
Horse with coat color that consists of large patches
color. Pinto patterns are visually and genetically distinct from the leopard complex spotting patterns characteristic of horse breeds such as the Appaloosa
Pinto_horse
Horse coat coloration
of a gray. This type of roaning only occurs in conjunction with the Leopard complex. Varnish roans are born with a dark base coat color, usually with some
Varnish_roan
Colored areas, usually white, on a horse that differ from the body color
dun as opposed to a buckskin or a bay. Leopard complex: Responsible for a variety of patterns, typically leopard-type spotting, and is most closely identified
Horse_markings
Coat color characterized by progressive depigmentation of the colored hairs of the coat
of Thoroughbreds. Gray also affects spotting patterns of pintos and leopard complex horses such as Appaloosas. Its effects wash out the contrast of the
Gray_horse
Allele responsible for several horse coat colors
genes underlying tobiano, dominant white, frame overo spotting and the Leopard complex common to the Appaloosa, the champagne gene does not affect the location
Champagne_gene
Animal with white markings on a darker coat
color variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Piebald
Dilution gene
do not. Prior to domestication of the horse, dun, non-dun1, and the leopard complex are thought to have been the wild type modifiers of the base colors
Dun_gene
Main battle tank family of German origin
The Kampfpanzer Leopard, subsequently Leopard 1 following the introduction of the successive Leopard 2, is a main battle tank designed by Porsche, and
Leopard_1
Genetics behind the equine coat color
thought to be much more recent, possibly from after domestication. Leopard complex patterns also predate domestication, having been found in horse remains
Equine_coat_color_genetics
Horse coat color
pink-skinned white markings. White markings and patterns such as pinto and leopard have no bearing on the underlying base coat color of the animal. Black
Black_horse
Coat coloring pattern in some animals
markings and ticking can occur on any brindle dog. Brindle is caused by a complex gene process and is technically a form of mosaicism, where some cells express
Brindle
Color of horses in the dun family
color variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Grullo
Spotted color pattern in horses
color variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Tobiano
Leopard subspecies
The African leopard (Panthera pardus pardus) is the nominate subspecies of the leopard, native to Africa. It is widely distributed in most of sub-Saharan
African_leopard
Hair coat color of horses
that do not have a category for pinto. Bay Leopards are horses that carry the leopard (Lp) gene or gene complex characteristic of the Appaloosa and other
Bay_(horse)
Coat color found in many animals
roans. A varnish roan is not a true roan; it is actually one of the leopard complex coat patterns associated with Appaloosa, Knabstrupper, Noriker horse
Roan_(color)
Pattern of irregular marks on plants or animals
but may also be a hereditary trait, as seen with the champagne and leopard complex genes in horses. "Mottles" can also refer to discoloration in processed
Mottle
Gene for several horse coat colors
E allele. This effect has also been identified and studied in the Leopard complex patterns. The SLC45A2 gene is best known in humans as being the location
Cream_gene
Equine coat color
color variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Buckskin_(horse)
Condition making it difficult or impossible to see in relatively low light
has been linked with the leopard complex color pattern since the 1970s. A 2008 study theorizes that both CSNB and leopard complex spotting patterns are linked
Nyctalopia
Leopard subspecies
Indochinese leopard (Panthera pardus delacouri) is a leopard subspecies native to mainland Southeast Asia and southern China. In Indochina, leopards are rare
Indochinese_leopard
Topics referred to by the same term
Sundaland islands Leopard (pattern), the spotting pattern characteristic of Leopards Leopard complex, the genetic allele responsible for leopard-spotting in
Leopard_(disambiguation)
Horse coat color
color variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Chestnut_(horse_color)
Species of wild cat
The clouded leopard (Neofelis nebulosa), also called mainland clouded leopard, is a wild cat that has large dusky-grey blotches and irregular spots and
Clouded_leopard
Hair coloration in some equine
color variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Primitive_markings
Colour
colour variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Isabelline_(colour)
Horse coat color and its genetics
patterns. The fewspot leopard pattern, however, can resemble white. Two factors influence the eventual appearance of a leopard complex coat: whether one copy
Dominant_white
American breed of horse
areas that were once dark. Spots can be any shape or size, except leopard complex patterning, which is characteristic of the Appaloosa, and located virtually
American_Paint_Horse
Leopard subspecies
The Indian leopard (Panthera pardus fusca) is a subspecies of the leopard (P. pardus). It is widely distributed on the Indian subcontinent. It is threatened
Indian_leopard
Colour pattern of horses
color variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Skewbald
2007 operating system version
Mac OS X Leopard (version 10.5) is the sixth major release of macOS, Apple's desktop and server operating system for Macintosh computers. Leopard was released
Mac_OS_X_Leopard
Coat trait found in some horses
color variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Pangaré
Group of colouration patterns of horses
color variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Overo
DNA profile of a horse
mapping of the horse genome was locating the mutation that creates the Leopard complex (Lp) spotting pattern seen in breeds such as the Appaloosa. Horses
Horse_genome
Lighter mane and tail on chestnut horses
color variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Flaxen_(color_variant)
Gene that lightens the coat colour of certain animals
color variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Dilution_gene
Color of horses' eyes
color variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Tiger_eye
Type of dilution gene
color variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Pearl_gene
Medical condition
involved. Two forms of CSNB can also affect horses, one linked to the leopard complex of equine coat colors and the other found in certain horse breeds.
Congenital stationary night blindness
Congenital_stationary_night_blindness
Hair coat color of horses
color variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Seal_brown_(horse)
Horse coat colors and patterns, such as white ticking or roaning
color variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Rabicano
colour variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Tricoloured_horse
Horse fur marking
color variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Leukotrichia
Type of spotted marking on horses
color variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Bend-Or_spots
Extinct subspecies of equines
from the Pleistocene and Holocene - those creating bay, black, and leopard complex are known from the wild horse population in Europe and were depicted
Tarpan
Horse coat color
color variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Smoky_black
Gene contributing to horse coat color
color variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Silver_dapple_gene
Trait characterized by black or darker hairs mixed into a horse's coat
color variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Sooty_horse
Horse coat colour
color variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Splashed_white
color variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Mushroom_gene
color variations (primarily UK English) Piebald Skewbald Tricoloured Leopard complex Varnish roan See also Appaloosa and Knabstrupper Roaning patterns Roan
Tovero
Color pattern in horses
testing. The genetics behind white patterns and markings in horses are complex. Two full siblings with the same genotype for a particular white spotting
Sabino_horse
Species of amphibian
pipiens formerly Rana pipiens, commonly known as the northern leopard frog, is a species of leopard frog from the true frog family, native to parts of Canada
Northern_leopard_frog
stockings or socks. Also used to refer to particularly flashy pinto or leopard spotting. cinch A wide flat strap made of mohair, reinforced felt, or an
Glossary_of_equestrian_terms
Topics referred to by the same term
darker-colored hairs Roan (horse), a horse coat color pattern Varnish roan, a leopard-complex horse coat color that looks similar to roan Roan antelope, an African
Roan
Horse breed originating in West Asia
Nonetheless, purebred Arabians today do not carry genes for pinto or Leopard complex ("Appaloosa") spotting patterns, except for sabino. Spotting or excess
Arabian_horse
Horse with spotting or markings from parents of solid color
crop-out or crop out is a horse with body spots, including pinto or leopard complex spotting, or "high white" horse markings, with a sire and dam who both
Cropout
Breed of horse
Association accepts horses of all colors and patterns, with the exception of leopard complex spotting, which is not accepted by the registry. There are three recognized
Quarab
Species of cartilaginous fish
The leopard whipray (Himantura leoparda) is a little-known species of stingray in the family Dasyatidae, found in the central Indo-West Pacific region
Leopard_whipray
Big cat crossbreed
five species of the genus Panthera: the tiger, lion, jaguar, leopard, and snow leopard. Most hybrids would not be perpetuated in the wild as the territories
Panthera_hybrid
American breed of horse
as "loud" Appaloosa coloring. The coloration includes the typical leopard complex characteristics of mottling around the eyes, muzzle and genitalia,
Pony_of_the_Americas
Chinese pony breed
coat is bay, chestnut, black, or sometimes gray or, even more rarely, leopard complex. White markings are possible on the head and lower limbs. Genetic studies
Datong_horse
Common name of several species of amphibian
Leopard frog is a generic name used to refer to various species in the true frog genus Lithobates. They all have similar coloration: brown or green with
Leopard_frog
Species of amphibian
The Chiricahua leopard frog (Lithobates chiricahuensis syn. Rana chiricahuensis) is a species of frog in the family Ranidae, the true frogs. It is native
Chiricahua_leopard_frog
Zoo in Washington, D.C., and Front Royal, Virginia
of summer 2025. In early 2026, plans were unveiled for an Arabian leopard complex that would replace the existing bison exhibit. Other animals in the
National Zoological Park (United States)
National_Zoological_Park_(United_States)
Mountain range in western India
restore the mountain range. The Sariska–Delhi leopard wildlife corridor or the Northern Aravalli leopard wildlife corridor is a 200 km long important biodiversity
Aravalli_Range
Georgian horse breed
chestnut as the most common (45%), followed by gray (28%). Pinto and leopard complex coats are rare, but possible. White markings are rare overall. With
Tushetian_horse
Plug-in hybrid full-size SUV
in China as the Fangchengbao Bao 8 (Chinese: 方程豹 豹8; lit. 'Formula leopard Leopard 8') is a plug-in hybrid full-size SUV manufactured by BYD Auto under
Denza_B8
Taiwanese/Irish wheeled fighting vehicle
The Clouded Leopard Eight-Wheeled Armored Vehicle (Chinese: 雲豹八輪甲車; pinyin: yúnbào bā lún jiǎ chē; lit. 'Cloud Leopard Eight-Wheeled Armored Vehicle')
Clouded Leopard Armored Vehicle
Clouded_Leopard_Armored_Vehicle
Species of amphibian
The Vegas Valley leopard frog (Lithobates fisheri), also known as the Las Vegas leopard frog, is an extinct species of frog. It once occurred in the Las
Vegas_Valley_leopard_frog
Species of amphibian
commonly known as the southern leopard frog, is a medium-sized anuran in the family Ranidae (the true frogs). The southern leopard frog is one of the 36 species
Southern_leopard_frog
Genus of amphibians
from the Early Miocene of Florida, and appear to belong to the leopard frog species complex. These species are recognised in the genus Lithobates: Lithobates
Lithobates
Breed of horse
crossed on an Appaloosa parent, so long as the ensuing foal also has leopard complex traits. Colorado Ranger: Similar to Appaloosa, has Arabian foundation
Part-Arabian
Rose-like markings on fur and skin
snow leopard (Panthera uncia) make it well adapted to life in cold climates with bare rocks and patchy snow. The rosette patterns in the leopard (P. pardus)
Rosette_(zoology)
Topics referred to by the same term
large white area of coloration in horses from the coat patterns of the leopard complex snow-capped manakin (Lepidothrix nattereri), a bird snowcap (Microchera
Snow_cap
2019 novel by Marlon James
Black Leopard, Red Wolf is a 2019 fantasy novel by Jamaican writer Marlon James. It is the first book of the Dark Star Trilogy. The novel draws on African
Black_Leopard,_Red_Wolf
Cave and archaeological site in France
study using the DNA of ancient horses, found that the leopard complex, which is involved in leopard spotting, was present, and concluded that the cave painters
Pech_Merle
of large mammals, including Indochinese tiger, Indochinese leopard, dhole, clouded leopard, sun bear, 10 species of primates (all five of the region's
Western_Forest_Complex
Species of amphibian
The plains leopard frog (Lithobates blairi) is a spotted frog found in North America. It is sometimes referred to as Blair's leopard frog, named after
Plains_leopard_frog
Film studio complex in Mumbai, India
The complex is surrounded by forested areas that are part of the national park, which serves as a habitat for wildlife, including leopards. Leopard sightings
Film_City,_Mumbai
Species of amphibian
The lowland leopard frog (Lithobates yavapaiensis) is a species of frog in the family Ranidae that is found in Mexico and the United States. Its natural
Lowland_leopard_frog
Hindu temple in Delhi, India
Nageshwar Mandir within the temple complex. Surroundings are an important biodiversity area within the Northern Aravalli leopard wildlife corridor stretching
Chhatarpur_Temple
2002 novel by Matthew Lynn (pen name James Harland)
his Estonian wife's mysterious disappearance and the powerful and complex Leopard Fund. Kirkus Reviews wrote, "Tension, pitifully lacking in the first
The_Month_of_the_Leopard
Species of amphibian
Rana pipiens complex: A biochemical evaluation. Systematic Zoology 32: 132–143. Hillis, D.M. & Frost, J.S. (1985): Three new species of leopard frogs (Rana
Showy_leopard_frog
Large feline of the genus Acinonyx
and leopard (Panthera pardus) as authors often confused the two; some considered "hunting leopards" an independent species, or equal to the leopard. In
Cheetah
Species of amphibian
Hillis, D.M.; Frost, J.S. (1985). "Three new species of leopard frogs (Rana pipiens complex) from the Mexican Plateau". Occ. Pap. Mus. Nat. Hist. Univ
Tlaloc's_leopard_frog
Japanese manga series by Paru Itagaki
Beast Complex (stylized in all caps) is a Japanese manga series written and illustrated by Paru Itagaki. It is a compilation of short stories set in the
Beast_Complex
Species of lizard
Anolis marmoratus, commonly known as the leopard anole, Guadeloupe anole, or Guadeloupean anole, is a species of anole that is endemic to the islands
Anolis_marmoratus
Street View "Zookeeper Down" December 29, 1994 Scottsbluff, Nebraska When a leopard attacks a zookeeper while she is trying to clean its cage, coworkers must
List_of_Rescue_911_episodes
American baseball player (born 2001)
Bloss spent his first professional season with the rookie-level Florida Complex League Astros and Fayetteville Woodpeckers, accumulating a 2.89 ERA in
Jake_Bloss
upcoming Mr. Universe contest. Bilko tricks Emma Ritzik into giving him her leopard skin coat to make a loincloth for Forbes. Bilko hires a woman (Lucille
List of The Phil Silvers Show episodes
List_of_The_Phil_Silvers_Show_episodes
LEOPARD COMPLEX
LEOPARD COMPLEX
Male
English
Variant spelling of English Leonard, LENARD means "lion-strong."
Boy/Male
Muslim
Little leopard
Surname or Lastname
English and Scottish
English and Scottish : patronymic from Leonard.
Boy/Male
Indian
Little leopard
Male
German
Variant spelling of German Leonhardt, LEONHARD means "lion-strong."
Surname or Lastname
English
English : from Middle English, Old French lepard ‘leopard’ (from Late Latin leopardus, a compound of leo ‘lion’ + pardus ‘panther’), probably applied as a nickname or as a habitational name for someone who lived at a house distinguished by the sign of a leopard.
Male
English
English form of French Léonard, LEONARD means "lion-strong."Â
Male
English
English form of French Léopold, LEOPOLD means "people-bold."
Male
Italian
Italian, Portuguese and Spanish form of German Leonhard, LEONARDO means "lion-strong."
Boy/Male
Muslim
Leopard
Male
English
Variant spelling of English Leonard, LENNARD means "lion-strong."
Surname or Lastname
English
English : variant spelling of Leopard.
Boy/Male
Indian
Leopard
Girl/Female
Muslim
Leopard
Girl/Female
Arabic, Muslim
Leopard
Surname or Lastname
English
English : variant of Liddiard.Revolutionary soldier William Ledyard was born at Groton, CT, in 1738, a descendant of John Ledyard who sailed from Bristol, England, and settled in CT. The celebrated traveler John Ledyard (1751–89) was William’s nephew and was also born in Groton.
Surname or Lastname
English and French (Léonard)
English and French (Léonard) : from a Germanic personal name composed of the elements leo ‘lion’ (a late addition to the vocabulary of Germanic name elements, taken from Latin) + hard ‘hardy’, ‘brave’, ‘strong’, which was taken to England by the Normans. A saint of this name, who is supposed to have lived in the 6th century, but about whom nothing is known except for a largely fictional life dating from half a millennium later, was popular throughout Europe in the early Middle Ages and was regarded as the patron of peasants and horses.Irish (Fermanagh) : adopted as an English equivalent of Gaelic Mac Giolla Fhionáin or of Langan.Americanized form of Italian Leonardo or cognate forms in other European languages.The French Léonard family were at Château Richer, Quebec, by 1698, having come from Maine, France.
Surname or Lastname
English
English : variant spelling of Leopard.
Boy/Male
Israeli
Leopard.
Girl/Female
Arabic
Female Leopard
LEOPARD COMPLEX
LEOPARD COMPLEX
Boy/Male
Hindu
Mine of nectar
Boy/Male
Australian, Gaelic, Irish
One who Sings Ballads
Girl/Female
Gujarati, Hindu, Indian, Kannada, Malayalam, Marathi, Sindhi, Tamil, Telugu
Understood and Accepted
Surname or Lastname
English
English : probably a habitational name from a lost or unidentified place; the forms Battershall and Battershill are also found.
Boy/Male
Tamil
Harendra | ஹரேநà¯à®¤à¯à®°
Lord Shiva, A tree
Girl/Female
Hindu, Indian
Good Friend; Soul Mate
Boy/Male
Hindu, Indian
Star
Boy/Male
Arthurian Legend American Latin Celtic English French Welsh
A knight.
Boy/Male
Australian, German, Latin, Scandinavian
Majestic; Staff of Gods; Meditation Staff
Boy/Male
Hindu
Sweet
LEOPARD COMPLEX
LEOPARD COMPLEX
LEOPARD COMPLEX
LEOPARD COMPLEX
LEOPARD COMPLEX
n.
Jeopardy.
n.
A leopard.
a.
Pertaining to, or in the direction of, the part or side toward which the wind blows; -- opposed to windward; as, a leeward berth; a leeward ship.
n.
A flowerlike color marking; as, the rosettes on the leopard.
n.
One who puts in jeopardy.
adv.
Toward the lee.
n.
A large, savage, carnivorous mammal (Felis leopardus). It is of a yellow or fawn color, with rings or roselike clusters of black spots along the back and sides. It is found in Southern Asia and Africa. By some the panther (Felis pardus) is regarded as a variety of leopard.
v. t.
To put in jeopardy; to expose to loss or injury; to imperil; to hazard.
v. t.
To expose to loss or injury; to risk; to jeopard.
n.
A tree, the top of which has been lopped off.
p. pr. & vb. n.
of Jeopard
n.
A leopard.
n.
To risk, or hazard; jeopard; to venture.
n.
A leopard; a panther.
n.
Exposure to death, loss, or injury; hazard; danger.
n.
The lee side; the lee.
v. t.
To jeopardize.
imp. & p. p.
of Jeopard