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Extinct order of trilobites
Proetida is an order of trilobite that lived from the Ordovician to the Permian. It was the last surviving order of trilobite, dying out in the Permian-Triassic
Proetida
Class of extinct, Paleozoic arthropods
Hangenberg/end-Devonian mass extinction, wiping out most trilobite diversity and leaving Proetida as the only surviving order. Their diversity moderately recovered during
Trilobite
Extinct order of trilobites
Asaphida, Proetida, Harpetida, and possibly Phacopida. These are now known as the "Librostome Orders". Trilobites of the orders Proetida, Harpetida,
Ptychopariida
System that relates geologic strata to time
sharks and ray-finned bony fish. Placoderms and graptolites die out. Proetida only group of trilobites. First freshwater molluscs and sharks. Arthropleura
Geologic_time_scale
Fourth period of the Paleozoic Era
placoderms, and nearly all trilobites save for a few species of the order Proetida. The subsequent end-Devonian extinction, which occurred at around 359 Ma
Devonian
Family of trilobites (fossil)
continued to recognize Phillipsiidae as a distinct and separate family within Proetida. The following genera are included, divided among six subfamilies: No subfamily
Phillipsiidae
Extinct genus of trilobites
Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Aulacopleuridae Genus: †Cyphaspis Burmeister 1843 Type species
Cyphaspis
Extinct order of trilobites
Librostoma. Phacopida was one of only two trilobite orders (along with the Proetida) to survive the Kellwasser event (Late Devonian extinction) at the Frasnian-Famennian
Phacopida
Order †Harpetida Order †Redlichiida Order †Lichida Order †Phacopida Order †Proetida Order †Ptychopariida Order †Isoxyida Order †Hymenocarina Order †Mollisoniida
List_of_arthropod_orders
Extinct superfamily of trilobite
Proetoidea is a superfamily of trilobites in the order Proetida. The Proetoidea are closely related to the Aulacopleuroidea. and are divided into three
Proetoidea
Geologic formation in Bolivia
Chacomurus C. confragosus Dalmanitidae Maurotarion M. legrandi Aulacopleuridae (Proetida) Bainella B. (Belenops) insolita Calmoniidae Deltacephalaspis D. (Deltacephalaspis)
Belén_Formation
member of Proetida belonging to the family Aulacopleuridae. Otarionella givetica Sp. nov Valid Flick & Flick Devonian Germany A member of Proetida belonging
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Geologic formation in Australia
Kettneraspis K. hollowayi Odontopleurida, Odontopleuridae Maurotarion M. euryceps Proetida, Aulacopleuridae Orygmatos O. yanyeani Phacopida, Phacopidae Raphiophorus
Melbourne_Formation
Subfamily of trilobites
Earth and Life. pp. 535–555. doi:10.1007/978-90-481-3428-1_16. ISBN 978-90-481-3427-4. "The Last Trilobites". trilobites.info. Proetida Fact Sheet v t e
Cummingellinae
Extinct genus of trilobites
Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Phillipsiidae Subfamily: †Archegoninae Genus: †Archegonus Burmeister
Archegonus
Geological formation in Western Australia
The Virgin Hills Formation is a geologic formation dating to the Late Devonian epoch, specifically the Frasnian and Famennian stages. It is located in
Virgin_Hills_Formation
Second of the big five mass extinctions
This left only two trilobites orders in the Famennian: Phacopida and Proetida. Trilobites which survived the Kellwasser event tended to prefer deep environments
Late_Devonian_mass_extinction
Ordovician Lagerstätte in France
Pharostomatidae. Proetidella? P.? sp. A trilobite belonging to the order Proetida. Selenopeltis S. binodosus A trilobite belonging to the family Odontopleuridae
Cabrières_Biota
Genus of phacopid trilobites
structure of their eyes Basse, Martin (Dec 2006). Eifel Trilobites vol. 4: Proetida 3, Phacopida 3. Goldschneck Verlag. ISBN 9783494014111. McKeller, Ryan;
Morocops
Extinct genus of trilobite
Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Phillipsiidae Subfamily: †Ditomopyginae Genus: †Ditomopyge Newell
Ditomopyge
Genus of trilobites
Fukushima Prefecture, Japan. It was originally described by Riuji Endo as Paladin yanagisawai. Proetida fact sheet [1] Endops in Paleobiology db [2] v t e
Endops
Extinct genus of trilobites
Kingdom: Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Hystricuridae Genus: †Etheridgaspis Kobayashi 1940
Etheridgaspis
Extinct order of arthropods
Asaphida Corynexochida Harpetida Lichida Nektaspida Odontopleurida Phacopida Proetida Ptychopariida Redlichiida Genera List of trilobites Geochronology Cambrian
Agnostida
Genus of trilobites
P. kussicum, P. anfensis, and P. netschaewi) Paraphillipsia vnweberi "PROETIDA FACT SHEET". trilobites.info. "Paraphillipsia". Mindat.org. "The Last Trilobites"
Paraphillipsia
Extinct order of trilobites
Asaphida Corynexochida Harpetida Lichida Nektaspida Odontopleurida Phacopida Proetida Ptychopariida Redlichiida Genera List of trilobites Geochronology Cambrian
Harpetida
Group of Devonian geologic formations in New South Wales, Australia
Calymenidae Otarion O. dabrowni Receptaculites Limestone Mbr, Taemas Proetida, Aulacopleuridae Phacops P. spedeni Receptaculites Limestone Mbr, Taemas
Murrumbidgee_Group
Extinct genus of trilobites
Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Proetidae Genus: †Aayemenaytcheia Lieberman, 1994 Species: †A
Aayemenaytcheia
Genus of arthropods (fossil)
classification Kingdom: Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Dimeropygidae Genus: †Dimeropygiella Ross, 1951
Dimeropygiella
Extinct genus of telephinid trilobite
some Ordovician Telephinidae from Argentina, and larval patterns in the Proetida (Trilobita)". Journal of Paleontology. 73 (2): 219–239. doi:10.1017/S0022336000027736
Telephina
Extinct family of trilobites
Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Superfamily: †Bathyuroidea Family: †Telephinidae Marek, 1952 Genera and
Telephinidae
Geologic formation in Oregon, US
pseudodraschei Stenoscisma cf. plicatum Tiramnia semiglobosa Yakovlevia transversa Proetida Phillipsiidae Cummingella (Cummingella) sp. Conocardiida Conocardiidae
Coyote_Butte_Limestone
Genus of trilobites
Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Phillipsiidae Subfamily: †Cummingellinae Genus: †Persiax Lerosey-Aubril
Persiax
Extinct subclass of trilobites
order Ptychopariida; and all post-Devonian trilobites being in the order Proetida, who are librostomes. Cummingella belisama, a Phillipsiid Proetid from
Librostoma
Extinct genus of trilobites
Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Aulacopleuridae Genus: †Aulacopleura Hawle & Corda, 1847 Type
Aulacopleura
Extinct species of trilobite
Asaphida Corynexochida Harpetida Lichida Nektaspida Odontopleurida Phacopida Proetida Ptychopariida Redlichiida Genera List of trilobites Geochronology Cambrian
Eldredgeops_rana
Arenig Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Genus: †Oenonella Fortey, 1980
Oenonella
Time interval during the Ordovician period
Pterygometopus Strotactinus Sycophantia Synhomalonotus Toletanaspis Xystocrania Proetida of the Floian Acidiphorus Bathyuriscops Benthamaspis Biolgina Bolbocephalus
Arenig
Extinct family of trilobites
Pärnaste; Adrian Popp & Robert M. Owens (2009). "Distribution of the order Proetida (Trilobita) in Baltoscandian Ordovician strata" (PDF). Estonian Journal
Proetidae
analyses. A study on the phylogenetic relationships of Devonian members of Proetida is published by Jordan-Burmeister (2025). Collantes & Pates (2025) revise
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Geologic formation in Bolivia
"synziphosurine" chelicerate Maurotarion M. dereimsi, M. racheboufi Aulacopleuridae (Proetida) Bainella B. (Belenops) insolita Calmoniidae Deltacephalaspis D. (Prestalia)
Icla_Formation
Mass extinction at the end of the Devonian
diversity and body size. The two remaining orders of trilobites, Phacopida and Proetida, were strongly affected. The order Phacopida completely died out during
Hangenberg_event
Extinct order of trilobites
Asaphida Corynexochida Harpetida Lichida Nektaspida Odontopleurida Phacopida Proetida Ptychopariida Redlichiida Genera List of trilobites Geochronology Cambrian
Corynexochida
Devonian geologic formation in Queensland, Australia
section (locality 3) Proetida, Tropidocoryphidae Aulacopleura A. sp. Mytton Formation Undifferentiated section (locality 3) Proetida, Aulacopleuridae Burgesina
Broken_River_Group
Extinct genus of trilobites
78: 9–29. Gibb, Stacey; Chatterton, Brian (2010-01-01). "Gerastos (Order Proetida; Class Trilobita) from the Lower to Middle Devonian of the southern Moroccan
Gerastos
Van Viersen & Müller Devonian (Emsian to Eifelian) Morocco A member of Proetida. The type species is "Cornuproetus (Diademaproetus)" antatlasius Alberti
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Extinct genus of trilobites
Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Proetidae Genus: †Liobole Richter and Richter, 1949 Species Liobole
Liobole
Extinct genus of trilobites
Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Bathyuridae Genus: †Aponileus Hu, 1963 Type species †Aponileus
Aponileus
Extinct genus of trilobites
Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Aulacopleuridae Genus: †Chamaeleoaspis Basse, 2010 Type species
Chamaeleoaspis
Extinct genus of trilobites
Asaphida Corynexochida Harpetida Lichida Nektaspida Odontopleurida Phacopida Proetida Ptychopariida Redlichiida Genera List of trilobites Geochronology Cambrian
Bumastus
Extinct genus of toernquistiid trilobite
(March 1998). "Ontogeny and systematics of Toernquistiidae (Trilobita, Proetida) from the Ordovician of the Argentine Precordillera". Journal of Paleontology
Lasarchopyge
classification Kingdom: Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Bathyuridae Genus: †Grinnellaspis Poulsen, 1948
Grinnellaspis
Genus of trilobites
Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Phillipsiidae Subfamily: †Cummingellinae Genus: †Cummingella Reed
Cummingella
Genus of bathyurid trilobite from the Early Ordovician
Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Bathyuridae Genus: †Veeversaspis Smith, and Allen, 2023 Type species
Veeversaspis
Extinct genus of trilobites
Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Proetidae Genus: †Wagnerispina Gandl, 1977 Species Wagnerispina
Wagnerispina
Genus of trilobites
Phillipsidae from the Takakura-yama Group (Permian), Abukuma Massif in Japan. Earth Science (Chikyu Kagaku) 26:19-25 [1] Proetida fact sheet [2] v t e
Nipponaspis
Genus of trilobites
Asaphida Corynexochida Harpetida Lichida Nektaspida Odontopleurida Phacopida Proetida Ptychopariida Redlichiida Genera List of trilobites Geochronology Cambrian
Dinea
Species of trilobite
Asaphida Corynexochida Harpetida Lichida Nektaspida Odontopleurida Phacopida Proetida Ptychopariida Redlichiida Genera List of trilobites Geochronology Cambrian
Phacops_fecundus
Extinct genus of bathyurid trilobites
classification Kingdom: Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Bathyuridae Genus: †Raymondites Sinclair, 1944
Raymondites
Extinct genus of trilobites
Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Proetidae Genus: †Helokybe Thomas & Narbonne, 1979 Species Helokybe
Helokybe
Genus of late trilobite
"The Last Trilobites". www.trilobites.info. III, Dr. Samuel M. Gon. "PROETIDA FACT SHEET". www.trilobites.info. Mychko, Eduard V. (2025-05-24). "Lopingian
Kathwaia
classification Kingdom: Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Bathyuridae Genus: †Eleutherocentrus Clark, 1935
Eleutherocentrus
Extinct genus of trilobite
Anisopyge is an extinct genus of trilobite belonging to the order Proetida and family Phillipsiidae. Specimens have been found in Permian beds in North
Anisopyge
Genus of trilobites
this stage, together with all other trilobites with the exception of some Proetida. It can be found in Australia, the Czech Republic, France, Germany, Iran
Trimerocephalus
Protected area in France
organisms and cnidarians. Identified trilobites (classified by orders): Proetida Family Proetidae Gerastos catervus (van Viersen, 2006) (previous names:
Vireux-Molhain National Nature Reserve
Vireux-Molhain_National_Nature_Reserve
Genus of trilobites
Tiryns, son of King Abas of Argo. Proetus is the type genus of the order Proetida, and of the family Proetidae. The genus became a wastebasket taxon that
Proetus_(trilobite)
Extinct genus of trilobites
Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Brachymetopidae Genus: †Cordania Clarke, 1892 Species C. buicki
Cordania
Geologic formation in China
P. changyangensis Pliomeridae Huaqiao Protarchaeogonus P? huaqiaoensis Proetida Rorringtoniidae Huaqiao Goniophrys G. subcylindricus Telephinidae Huaqiao
Fenxiang_Formation
Genus of trilobites
Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Rorringtoniidae Genus: †Cyamella Owens in Owens & Hammann, 1990
Cyamella_(trilobite)
Extinct genus of trilobites
classification Kingdom: Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Bathyuridae Genus: †Ceratopeltis Poulsen, 1937
Ceratopeltis
Extinct genus of trilobites
Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Proetidae Genus: †Gitarra Gandl, 1968 Species G. leonensis Romano
Gitarra
Extinct genus of trilobites
Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Telephinidae Genus: †Carolinites Kobayashi, 1940 Type species
Carolinites
Arenig Scientific classification Kingdom: Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Genus: †Biolgina Maksimova, 1955
Biolgina
Genus of trilobites
Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Phillipsiidae Subfamily: †Ditomopyginae Genus: †Triproetus Steininger
Triproetus
Extinct genus of trilobites
classification Kingdom: Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Proetidae Genus: †Dechenella Kayser, 1880
Dechenella
classification Kingdom: Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Bathyuridae Genus: †Hadrohybus Raymond, 1925
Hadrohybus
Extinct genus of trilobites
Kingdom: Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Tropidocoryphidae Genus: †Diademaproetus Alberti, 1964
Diademaproetus
Extinct genus of trilobites
Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Tropidocoryphidae Genus: †Cornuproetus Richter & Richter, 1919
Cornuproetus
classification Kingdom: Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Bathyuridae Genus: †Lutesvillia Cullison, 1944
Lutesvillia
classification Kingdom: Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Bathyuridae Genus: †Rananasus Cullison, 1944
Rananasus
6–11. David J. Holloway (2013). "The trilobite Cirriticeps gen. nov. (Proetida, Tropidocoryphidae) from the Silurian of New South Wales". Memoirs of the
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Genus of trilobites
Australia (i) Australokaskia (Australokaskia), Planokaskia, Malchi n. gen." Geologica et Palaeontologica 28 (1994): 79-101. Proetida fact sheet [1] v t e
Malchi
classification Kingdom: Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Bathyuridae Genus: †Benthamaspis Poulsen, 1947
Benthamaspis
classification Kingdom: Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Dimeropygidae Genus: †Celmus Angelin, 1854
Celmus
Genus of proetid trilobite belonging to the family Phillipsiidae
family Phillipsidae from the Takakura-yama Group (Permian), Abukuma Massif in Japan. Earth Science (Chikyu Kagaku) 26:19-25 Proetida fact sheet [1] v t e
Thaiaspis
Extinct genus of trilobites
classification Kingdom: Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Proetidae Genus: †Eocyphinium Reed, 1942
Eocyphinium
Van Viersen, van Rossum & Prescher Middle Devonian Belgium A member of Proetida. Australoscutellum iota sp nov Valid Holloway and Lane Silurian Australia
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moei Sp. nov Valid Flick & Flick Devonian (Eifelian) Germany A member of Proetida belonging to the family Aulacopleuridae and the subfamily Otarioninae.
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Genus of trilobites
Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Bathyuridae Genus: †Bolbocephalus Whitfield, 1890 Type species
Bolbocephalus
Extinct genus of trilobites
Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Phillipsiidae Subfamily: †Ditomopyginae Genus: †Pseudophillipsia
Pseudophillipsia
Extinct genus of trilobites
Hedstroemia is a genus of trilobites in the order Proetida known from the Silurian period of Europe, Asia, and North America. The genus Hedstroemia was
Hedstroemia_(trilobite)
nördlichen Schelf des Rheia-Ozeans (Mitteldevon: Givetium; Trilobita: Proetida; Rhenoherzynikum)". Dortmunder Beiträge zur Landeskunde - naturwissenschaftliche
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Extinct genus of trilobite
Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Phillipsiidae Subfamily: †Ditomopyginae Genus: †Ameura Weller
Ameura
Canadian botanist and geologist (1837–1923)
Asaphida Corynexochida Harpetida Lichida Nektaspida Odontopleurida Phacopida Proetida Ptychopariida Redlichiida Genera List of trilobites Geochronology Cambrian
George_Frederick_Matthew
Buchiproetus? exconcha Sp. nov Valid Flick Devonian Germany A member of Proetida belonging to the family Tropidocoryphidae. Buenellus chilhoweensis Sp.
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Genus of trilobites
present in immature individuals). Weller, J. Marvin. "Permian trilobite genera." Journal of Paleontology (1944): 320-327. Proetida fact sheet [1] v t e
Vidria
Extinct family of trilobites
Tropidocoryphidae is an extinct family of trilobites in the order Proetida. These 44 genera belong to the family Tropidocoryphidae: Aidynsaia Owens & Ivanova
Tropidocoryphidae
Belgium A member of Proetida. Astycoryphe tureki Sp. nov Valid Mergl & Budil Devonian (Eifelian) Czech Republic A member of Proetida. Bigotina kelleri Sp
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Extinct genus of trilobites
classification Kingdom: Animalia Phylum: Arthropoda Clade: †Artiopoda Class: †Trilobita Order: †Proetida Family: †Proetidae Genus: †Paladin Weller, 1936
Paladin_(trilobite)
member of Proetida belonging to the family Aulacopleuridae. Cyphaspides speculator Sp. nov Valid Alberti Devonian Germany A member of Proetida belonging
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