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Clade of reptiles
Testudinata is the group of all tetrapods with a true turtle shell. It includes both modern turtles (Testudines) and many of their extinct, shelled relatives
Testudinata
Species of barnacle
Lepas testudinata is a species of goose barnacle in the family Lepadidae. First observed in 1834, Lepas testudinata has undergone several reclassifications
Lepas_testudinata
Clade of reptiles
Pantestudines †Eosauropterygia †Placodontia †Sinosaurosphargis †Odontochelys Testudinata †Proganochelys Testudines Archosauromorpha †Choristodera †Prolacertiformes
Sauria
Clade of tetrapods
Weigeltisauridae † Order Younginiformes † (Infraclass) Clade Neodiapsida Order Testudinata Suborder Testudines – turtles Infraclass Lepidosauromorpha Unnamed infrasubclass
Amniote
Clade of reptiles
stem-turtles). Pantestudines with a complete shell are placed in the clade Testudinata. The phylogenetic placement of turtles relative to other reptiles has
Pantestudines
Species of moth
Dichomeris testudinata is a moth in the family Gelechiidae. It was described by Edward Meyrick in 1934. It is found in Guangdong, China. Beccaloni, G
Dichomeris_testudinata
Clade of reptiles
Pan-Testudines/ †Eosauropterygia †Placodontia †Sinosaurosphargis †Odontochelys Testudinata †Proganochelys Testudines Pantestudines Pan‑Archosauria †Choristodera
Archosauromorpha
Geological formation in Germany and Switzerland
The Trossingen Formation, formerly the Knollenmergel (also known as the Arnstadt Formation or Feuerletten Formation), is a geological formation in Germany
Trossingen_Formation
Group of Mesozoic aquatic reptiles
Pan-Testudines/ †Eosauropterygia †Placodontia †Sinosaurosphargis †Odontochelys Testudinata †Proganochelys Testudines Pantestudines Archosauromorpha s. l. Sauria
Sauropterygia
Taxonomic clade
Pan-Testudines/ †Eosauropterygia †Placodontia †Sinosaurosphargis †Odontochelys Testudinata †Proganochelys Testudines (turtles) Pantestudines Pan-Archosauria †Choristodera
Sauropsida
Class of animals
Pan-Testudines/ †Eosauropterygia †Placodontia †Sinosaurosphargis †Odontochelys Testudinata †Proganochelys Testudines (turtles) Pantestudines Pan-Archosauria †Choristodera
Reptile
Extinct clade of turtles
Paracryptodira is an extinct group of reptiles in the clade Testudinata (which contains modern turtles and their extinct relatives), known from the Jurassic
Paracryptodira
Family of turtles
John Edward (1869). "Notes on the families and genera of tortoises (Testudinata), and on the characters afforded by the study of their skulls". Proceedings
Chelydridae
Geological group in the Middle Atlas of Morocco
Genus Species Stratigraphic position Material Notes Images Testudinata indet. Indeterminate Tamguert n'Tarit Tizi n'Juillerh Very numerous debris: escutcheons
El_Mers_Group
Cretaceous marine turtle and the largest turtle ever discovered
"Reassessment of the Phylogenetic Interrelationships of Basal Turtles (Testudinata)". Journal of Systematic Palaeontology. 10 (1): 35. doi:10.1080/14772019
Archelon
Geologic formation in Poland and Germany
Material Notes Images Chelonipus C. isp. Sołtyków Footprints Turtle Tracks Testudinata indet. Indeterminate Sołtyków Carapace Fragments Unidentified Turtle
Zagaje_Formation
English zoologist, surgeon and writer
turtle Plestiodon lynxe bellii, a subspecies of lizard A Monograph of the Testudinata. 1832–1836. – summarizes all the world's turtles, living and extinct
Thomas_Bell_(zoologist)
Suborder of reptiles
"Reassessment of the phylogenetic interrelationships of basal turtles (Testudinata)". Journal of Systematic Palaeontology. 10 (1): 3–45. doi:10.1080/14772019
Cryptodira
Clade of turtles
less than Testudinata. The clade is known from the Early Jurassic to the Present, and contains all Jurassic representatives of Testudinata aside from
Mesochelydia
Order of reptiles with a shell
extinct relatives with a complete shell are classified within the clade Testudinata. The most recent common ancestor of living turtles, corresponding to
Turtle
This is an overview of the paleofauna of the Eocene Messel Formation as explored by the Messel Pit excavations in Germany. A former quarry and current
Paleofauna of the Messel Formation
Paleofauna_of_the_Messel_Formation
Topic in reptilian phylogenetics
Archelosauria). Below are many of the possible classifications of Testudines and Testudinata: Below is a cladogram of living testudines found by Thomson and Shaffer
Turtle_classification
Geological Formation in Baden-Württemberg, Germany
The Nusplingen Limestone (German: Nusplingen Plattenkalk) is a geological formation in Baden-Württemberg, Germany. It preserves fossils dating to the Kimmeridgian
Nusplingen_Limestone
Jurassic-era geologic formation in Switzerland
"A new species of the large-headed coastal marine turtle Solnhofia (Testudinata, Thalassochelydia) from the Late Jurassic of NW Switzerland". PeerJ.
Reuchenette_Formation
Geological formation in India
The Kota Formation is a geological formation in India. The age of the Kota Formation is uncertain; it is commonly considered to date to the Early Jurassic
Kota_Formation
List of reptile genera lists the vertebrate class of reptiles by living genus, spanning two subclasses. Turtles are reptiles of the order Testudines characterized
List_of_reptile_genera
British mineralogist, botanist, and illustrator
fathered seven children with her. Terrapene clausa from Monograph of the Testudinata (lithograph by Edward Lear) Sowerby died in Kilburn, London in 1871.
James_De_Carle_Sowerby
Extinct family of turtles
and evolution of the bizarre clade of horned turtles (Meiolaniidae, Testudinata)". Zoological Journal of the Linnean Society. 174 (3): 519–548. doi:10
Meiolaniidae
The following is a list of terrestrial vertebrate fauna of Mount Tây Côn Lĩnh, western Vị Xuyên District, Hà Giang Province, Vietnam, adapted from Nguyen
List_of_fauna_of_Hà_Giang
British scientific illustrator (1798–1873)
pale-throated sloth's bone structures were published in A Monograph of the Testudinata and Transactions of the Zoological Society of London. Bell was born to
Jane_S._Bell
Extinct genus of reptiles
Pantestudines †Eorhynchochelys †Eunotosaurus? †Odontochelys †Pappochelys Testudinata †Priscochelys? †Proganochelyidae Proganochelys Ypomonetikochelys †Proterochersidae
Odontochelys
British zoologist (1800–1875)
of Shield Reptiles in the Collection of the British Museum – Part 1, Testudinata (Tortoises). 1860-10 : "On the arrangement of the land pulmoniferous
John_Edward_Gray
Animalia Phylum: Chordata Class: Reptilia Clade: Pantestudines Clade: Testudinata Clade: †Paracryptodira Family: †Compsemydidae Genus: †Tongemys Joyce
Tongemys
Extinct genus of stem-turtle
Bartassochersis was recovered as a stem-turtle (early-diverging member of Testudinata outside of the turtle crown group, Testudines), within the family Proterochersidae
Bartassochersis
Clade of reptile-like animals
Microsauria, Diadectomorpha, Procolophonia, Pareiasauria, Captorhinidia, Testudinata. Michael Benton (2000, 2004) made it the sister-clade to Lepospondyli
Reptiliomorpha
Extinct genus of turtles
Animalia Phylum: Chordata Class: Reptilia Clade: Pantestudines Clade: Testudinata Family: †Sinemydidae Genus: †Sinemys Wiman, 1930 Species S. lens Wiman
Sinemys
Extinct genus of turtles
Animalia Phylum: Chordata Class: Reptilia Clade: Pantestudines Clade: Testudinata Clade: †Meiolaniformes Genus: †Patagoniaemys Sterli & de la Fuente, 2011
Patagoniaemys
Family of turtles
(2000). "On the identification of the holotype of Chelodina oblonga (Testudinata: Chelidae) with a discussion of the taxonomic implications". Chelonian
Chelidae
Species of turtle
of Shield Reptiles in the Collection of the British Museum. Part I. Testudinata (Tortoises). London: Trustees of the British Museum. (Taylor and Francis
Eastern_long-necked_turtle
Family of turtles
"Reassessment of the phylogenetic interrelationships of basal turtles (Testudinata)". Journal of Systematic Palaeontology. 10 (1): 3–45. doi:10.1080/14772019
Dermochelyidae
Animalia Phylum: Chordata Class: Reptilia Clade: Pantestudines Clade: Testudinata Clade: †Thalassochelydia Family: †Plesiochelyidae Genus: †Globochelus
Globochelus
Extinct genus of turtle
Szczygielski, Tomasz; Dróżdż, Dawid (27 March 2026). "Triassic turtles ( Testudinata : Proterochersidae and Proganochelyidae ) of Greenland". Zoological Journal
Ypomonetikochelys
Second period of the Mesozoic Era
otherwise known from the Cretaceous and Cenozoic. Stem-group turtles (Testudinata) diversified during the Jurassic. Jurassic stem-turtles belong to two
Jurassic
Species of turtle
432 pp. Baur, G. 1893. Notes on the classification and taxonomy of the Testudinata. Proc. Amer. Philos. Soc. 31: 210–225. Boulenger, G.A. 1889. Catalogue
Malayan_softshell_turtle
Family of land turtles
of Shield Reptiles in the Collection of the British Museum. Part I. Testudinata (Tortoises)". London: British Museum, 28 pp. Fitzinger, Leopold J. (1835)
Tortoise
Geological formation in China
known from China and an example of one of the earliest known globally Testudinata indet. Indeterminate Houchiatsun, sichuan province Weiyuan, Szechuan
Ziliujing_Formation
Genus of barnacles
pectinata Spengler, 1793 Lepas tanakai Karasawa & Kobayashi, 2025 Lepas testudinata Aurivillius, 1894 Images of Lepas A detached holdfast of Durvillaea antarctica
Lepas
Extinct genus of turtles
skull, for example, has an edentulous beak typical of all members of Testudinata. However, the thorax region is markedly different from Pappochelys and
Eorhynchochelys
Species of turtle
Underwater nesting by the tropical freshwater turtle, Chelodina rugosa (Testudinata: Chelidae). Australian Journal of Zoology 41:47-52. doi:10.1071/zo9930047
Northern_snake-necked_turtle
Species of turtle
of Shield Reptiles in the Collection of the British Museum. Part I. Testudinata (Tortoises). London: British Museum. (Taylor and Francis, printers).
Blanding's_turtle
Extinct genus of turtles
conforms to a semi-aquatic mode of life. Proganochelys belongs to the group Testudinata, which consists of all extant turtles and several taxa of extinct kin
Proganochelys
Clade of reptiles
be one of three groups of Pholidota (reptiles), the other two being Testudinata (turtles) and Squamata (lizards and snakes). Loricata was an early name
Loricata
Extinct genus of turtles
Animalia Phylum: Chordata Class: Reptilia Clade: Pantestudines Clade: Testudinata Family: †Sinemydidae Genus: †Xiaochelys Zhou and Rabi, 2015 Type species
Xiaochelys
Monotypic species of reptile
and evolution of the bizarre clade of horned turtles (Meiolaniidae, Testudinata)". Zoological Journal of the Linnean Society. 174 (3): 519–548. doi:10
Gaffneylania
Family of turtles
Gray, John Edward. 1869. Notes on the families and genera of tortoises (Testudinata), and on the characters afforded by the study of their skulls. Proceedings
Geoemydidae
Extinct genus of reptiles
Pantestudines †Eorhynchochelys †Eunotosaurus? †Odontochelys †Pappochelys Testudinata †Priscochelys? †Proganochelyidae Proganochelys Ypomonetikochelys †Proterochersidae
Pappochelys
Plioplatecarpinae indet. Pteranodontidae indet. Squalidae indet. Teleostei indet. Testudinata indet. Tylosaurinae indet. ?Thoracosaurinae indet. List of pterosaur-bearing
Rybushka_Formation
Extinct genus of pleurosternid turtle
Philosophical Society 17(100):193-196 O. C. Marsh. 1890. Notice of some extinct Testudinata. The American Journal of Science and Arts, series 3 40:177-179 O. P.
Glyptops
Former group of reptiles
Pantestudines? †Sauropterygia †Eosauropterygia †Sinosaurosphargis †Placodontia †Eunotosaurus †Pappochelys †Odontochelys Testudinata †Proganochelys Testudines
Ankylopoda
Human protein-coding gene
Squamata 320 331 49% 63% Chelonia mydas Green Turtle XP_007069233.1 Testudinata 320 345 49% 61% Alligator mississippiensis American alligator XP_019353135
C19orf67
Extinct genus of turtles
Animalia Phylum: Chordata Class: Reptilia Clade: Pantestudines Clade: Testudinata Clade: †Thalassochelydia Family: †Thalassemydidae Genus: †Palaeomedusa
Palaeomedusa
Extinct clade of reptiles
(Argentina) on the evolution and palaeo-biogeography of Meiolaniformes (Testudinata, new taxon name)". Journal of Systematic Palaeontology. 11 (7): 835–852
Meiolaniformes
Extinct genus of turtles
"A new species of the large-headed coastal marine turtle Solnhofia (Testudinata, Thalassochelydia) from the Late Jurassic of NW Switzerland". PeerJ.
Solnhofia
Extinct genus of turtles
"Reassessment of the phylogenetic interrelationships of basal turtles (Testudinata)". Journal of Systematic Palaeontology. 10: 3–45. doi:10.1080/14772019
Kayentachelys
Extinct genus of turtles
Animalia Phylum: Chordata Class: Reptilia Clade: Pantestudines Clade: Testudinata Clade: †Paracryptodira Family: †Pleurosternidae Genus: †Selenemys Pérez-García
Selenemys
Extinct turtle species
Animalia Phylum: Chordata Class: Reptilia Clade: Pantestudines Clade: Testudinata Clade: †Thalassochelydia Genus: †Caesaria Spicher et al., 2026 Species:
Caesaria_templaria
Genus of turtles
of Shield Reptiles in the Collection of the British Museum. Part I. Testudinata (Tortoises). London: British Museum, 120 pp. Lindholm, Wassili A. (1931)
Mauremys
Genus of turtles
1834 by English zoologist Thomas Bell in his work A Monograph of the Testudinata. Cyclemys and closely related genera (Cuora, Pyxidea, and Notochelys)
Cyclemys
Extinct family of turtles
zoologist George Albert Boulenger published his classification of the Testudinata within the 9th edition of the Encyclopædia Britannica. The genus Protostega
Protostegidae
Extinct genus of compsemyid turtle
2026). "Re-Evaluation of Compsemys and North American Compsemydids (Testudinata)". Taxonomy. 6 (3): 48. doi:10.3390/taxonomy6030048. Martin Jehle (April
Compsemys
Extinct genus of turtles
Animalia Phylum: Chordata Class: Reptilia Clade: Pantestudines Clade: Testudinata Family: †Macrobaenidae Genus: †Yakemys Tong et al., 2021 Type species
Yakemys
Genus of mites
Hydrachna tchadensis Smit, 1994 Hydrachna tenuissima Viets, 1935 Hydrachna testudinata Cook Hydrachna trilobata Viets & K. Hydrachna triscutata Lundblad, 1947
Hydrachna
Geologic formation in New Mexico, USA
Taxon Locality Material Notes Testudinata Elephant Butte (upper) Crocodylia Compsemys Index fossils suggesting a Lancian age Bothremydidae
Hall_Lake_Formation
Fossil flora and fauna in the Posidonia Shale, Germany
almost complete skeleton Related to Eileanchelys and Heckerochelys. Testudinata indet. Indeterminate Altdorf Banz Hondelange Shells? and Isolated plastrons
Paleobiota of the Posidonia Shale
Paleobiota_of_the_Posidonia_Shale
Species of turtle
sous-espèce originaire du Paraguay, Phrynops tuberculatus vanderhaegei (Testudinata – Pleurodira – Chelidae) ". Bulletin de la Société Zoologique de France
Vanderhaege's toad-headed turtle
Vanderhaege's_toad-headed_turtle
Extinct genus of turtles
Animalia Phylum: Chordata Class: Reptilia Clade: Pantestudines Clade: Testudinata Clade: Rhaptochelydia Family: †Australochelyidae Genus: †Palaeochersis
Palaeochersis
Extinct genus of turtles
meaning Chinle turtle) is an extinct genus of stem-turtle belonging to Testudinata. It lived in the Norian age of the Late Triassic and is the oldest turtle
Chinlechelys
Species of turtle
of Shield Reptiles in the Collection of the British Museum. Part I. Testudinata (Tortoises). London: British Museum, 79 pp. [Published Mar 1856]. Uwe
Mesoclemmys_raniceps
Extinct genus of turtles
Animalia Phylum: Chordata Class: Reptilia Clade: Pantestudines Clade: Testudinata Clade: †Paracryptodira Family: †Pleurosternidae Genus: †Pleurosternon
Pleurosternon
Extinct genus of turtles
Animalia Phylum: Chordata Class: Reptilia Clade: Pantestudines Clade: Testudinata Clade: †Paracryptodira Family: †Pleurosternidae Genus: †Toremys Pérez-García
Toremys
Extinct genus of turtle
Animalia Phylum: Chordata Class: Reptilia Clade: Pantestudines Clade: Testudinata Clade: †Paracryptodira Family: †Baenidae Genus: †Lakotemys Joyce et al
Lakotemys
Species of turtle
ecología y distribución geográfica de Phrynops (Batrachemys) dahli (Testudinata, Pleurodira, Chelidae) ". Caldasia 9 (45): 467–489. (in Spanish). https://web
Dahl's_toad-headed_turtle
Genus of turtles
of Shield Reptiles in the Collection of the British Museum. Part I. Testudinata (Tortoises). The Trustees (British Museum of natural History), London
Pseudemys
Mesozoic geologic formation in Italy
be recognized. Overall about fifteen finds referable to sea turtles Testudinata indet. Indeterminate Villaggio del Pescatore Križ, southwest of Tomaj
Lipica_Formation
turtles of France: the first European Rhaetian stem turtle taxon (clades Testudinata, Proterochersidae)". Swiss Journal of Palaeontology. 145: 851–879. doi:10
2026_in_reptile_paleontology
Extinct genus of baenid turtles
Rollot, Y. (2023). "A description of a Denazinemys nodosa specimen (Testudinata, Baenidae) from the Late Cretaceous Kaiparowits Formation of southern
Denazinemys
Extinct genus of turtles
and evolution of the bizarre clade of horned turtles (Meiolaniidae, Testudinata)". Zoological Journal of the Linnean Society. 174 (3): 519–548. doi:10
Warkalania
Fossiliferous beds in North Africa
Ägyptens. II. Wirbeltierreste der Baharije-Stufe (unterstes Cenoman). 14. Testudinata". Abhandlungen der Bayerischen Akademie der Wissenschaften, Mathematisch
Bahariya_Formation
Genus of turtles
of Shield Reptiles in the Collection of the British Museum. Part I. Testudinata (Tortoises). With Figures of the Skulls of 36 Genera. London: Trustees
Sacalia
Clark, Brendan A. (2026). Paleobiology of a New "Macrobaenid" Turtle (Testudinata: Pan-Cryptodira) from the Blackleaf Formation, Montana (MSc thesis).
List of informally named Mesozoic reptiles
List_of_informally_named_Mesozoic_reptiles
Extinct family of turtles
Animalia Phylum: Chordata Class: Reptilia Clade: Pantestudines Clade: Testudinata Clade: †Paracryptodira Family: †Compsemydidae Pérez-García et al., 2015
Compsemydidae
Indian zoologist (1943–2019)
a Parasite of Buccal Cavity of Trachemys callirostris callirostris (Testudinata: Emydidae) from Colombia". The Journal of Parasitology. 95 (4): 850–854
K._C._Pandey
Extinct genus of turtles
Remarks on the Patagonian Horned Turtle Niolamia argentina Ameghino, 1899 (Testudinata, Meiolaniidae)" (PDF). Journal of Vertebrate Paleontology. 31 (6): 1210–1229
Niolamia
Extinct genus of turtles
Animalia Phylum: Chordata Class: Reptilia Clade: Pantestudines Clade: Testudinata Clade: †Thalassochelydia Genus: †Jurassichelon Pérez García, 2015 Type
Jurassichelon
Extinct clade of turtles
Animalia Phylum: Chordata Class: Reptilia Clade: Pantestudines Clade: Testudinata Clade: Mesochelydia Clade: Perichelydia Clade: †Thalassochelydia Anquetin
Thalassochelydia
Species of turtle
species pair of freshwater turtles, Elseya purvisi and Elseya georgesi (Testudinata: Chelidae), from eastern Australia". Chelonian Conservation and Biology
Bellinger River snapping turtle
Bellinger_River_snapping_turtle
Genus of fossil turtle
Animalia Phylum: Chordata Class: Reptilia Clade: Pantestudines Clade: Testudinata Family: †Macrobaenidae Genus: †Judithemys Parham & Hutchison, 2003 Type
Judithemys
Species of turtle
Zhao, "Er-mi"; Zhou, Ting; Ye, Ping (1990). "A new Chinese box turtle (Testudinata: Emydidae) — Cuora zhoui ". pp. 213–216. In: Zhao, "Ermi" (editor) (1990)
Zhou's_box_turtle
Extinct genus of turtles
Cyrtura is a dubious genus of extinct Testudinata from the Late Jurassic (early Tithonian) of the Solnhofen Formation of Bavaria, Germany. Cyrtura was
Cyrtura
Extinct genus of baenid turtle
Animalia Phylum: Chordata Class: Reptilia Clade: Pantestudines Clade: Testudinata Clade: †Paracryptodira Family: †Baenidae (unranked): †Eubaeninae Genus:
Boremys
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