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Extinct species of carnivore
bears. Ursus deningeri is widely thought to have evolved from Ursus etruscus (which is also thought to be the ancestor of the brown bear Ursus arctos)
Ursus_deningeri
Extinct species of carnivore
million years ago. U. deningeri is known from remains spanning from Europe to China. The transition between Ursus deningeri and Ursus spelaeus is often given
Cave_bear
Genus of bears
†Ursus dolinensis (Garcia & Arsuaga, 2001) †Ursus etruscus Cuvier, 1823 †Ursus minimus (Devèze & Bouillet, 1827) Cave bear complex: †Ursus deningeri Richenau
Ursus_(mammal)
Extinct subspecies of carnivore
these dimensions. Polar bear Ursus kudarensis Steppe brown bear Kurtén, B. (1964). "The evolution of the polar bear, Ursus maritimus Phipps". Acta Zoologica
Ursus_maritimus_tyrannus
Pre-Neanderthal population in Spain
complex at Atapuerca, and may have been a natural trap for the cave bear Ursus deningeri in particular. The Sima de los Huesos hominins may have been deposited
Sima_de_los_Huesos_hominins
Species in mammal family Ursidae
abstrusus (4.9–3.6 Mya) Genus Ursus U. arctos priscus (steppe brown bear) U. arvernensis (3.2–2.5 Mya) U. deningeri (0.79–0.12 Mya) (Deninger's bear)
List_of_ursids
Subfamily of bears
Ursus] – including †Ursus pyrenaicus (Depéret, 1892) and †Ursus sackdillingensis Heller, 1955 cave bears [= subgenus Spelearctos]: †Ursus deningeri Richenau
Ursinae
Extinct genus of bears
Ursini Ursus boeckhi Ursus thibetanus Ursus ursinus Ursus malayanus Ursus minimus Ursus americanus Ursus etruscus Ursus deningeri Ursus spelaeus Ursus arctos
Aurorarctos
Archaeological site in West Sussex, England
Hoxnian interglacial (MIS 11) or later interglacials. These include Ursus deningeri (an early member of the cave bear lineage), members of the giant deer
Boxgrove_Palaeolithic_site
Extinct species of carnivore
continental environments with cold and arid climate. Ursus ingressus and Ursus spelaeus evolved from Ursus deningeri and probably diverged between 173,000 and 414
Ursus_ingressus
Archaeological site in northern Spain, rich in human fossils
(also classified as early Neanderthals). Associated finds include Ursus deningeri fossils and a hand axe called Excalibur. It has received a surprisingly
Archaeological site of Atapuerca
Archaeological_site_of_Atapuerca
Extinct species of carnivore
of the living brown bear (Ursus arctos) and the extinct cave bears. The Etruscan bear appears to have originated from Ursus minimus and gave rise to the
Ursus_etruscus
Method of archaeological study
reported in 2013 the reconstructed mitochondrial genome of a bear, Ursus deningeri, more than 300,000 years old, proving that authentic ancient DNA can
Ancient_DNA
Extinct species of carnivore
bears (Ursus arctos) and noticeably gracile features of the cranial and mandibular bones, that to some extent resemble those of Ursus arctos and Ursus etruscus
Ursus_dolinensis
Cave and archeological site in Petralona, Chalkidiki, Greece
meles atavus ? (Kormos) (primitive European badger) † Ursus stehlini ? (Kretzoi) † Ursus deningeri Reichenau † Crocuta petralonae Kurten † Pachycrocuta
Petralona_Cave
Extinct species of carnivore
Ursus minimus (the Auvergne bear) is an extinct species of bear, endemic to Europe during the Pliocene and Pleistocene, living from 5.3 to 1.8 Mya, existing
Ursus_minimus
Extinct genus of bears
†Aurorarctos †Protarctos †Helarctos sinomalayanus †Melursus theobaldi Ursus †U. deningeri †U. dolinensis †U. etruscus †U. ingressus †U. kanivetz †U. kudarensis
Arctotherium
birds from 23 families, 51 genera and 68 species. Earlier (1943) were Ursus deningeri discovered, and later rich spectrum of coastal animal fossils such
Stránská_skála
Extinct genus of bears
Arctodus should not be mistaken for the similarly large Eurasian cave bear (Ursus spelaeus). As an ursine, the Eurasian cave bear last shared a common ancestor
Arctodus
Geological formation in Norfolk, England
"Cranial and mandibular morphology of Middle Pleistocene cave bears ( Ursus deningeri ): implications for diet and evolution". Historical Biology. 31 (4):
Cromer_Forest_Bed
free from inhibitory cascade constraints at least twice (Ursus minimus and Ursus deningeri) caused by dietary adaptations". Boreas. 55 (2) bor.70044
2025_in_paleomammalogy
Extinct genus of bears
†Aurorarctos †Protarctos †Helarctos sinomalayanus †Melursus theobaldi Ursus †U. deningeri †U. dolinensis †U. etruscus †U. ingressus †U. kanivetz †U. kudarensis
Ailurarctos
Extinct genus of bears
†Aurorarctos †Protarctos †Helarctos sinomalayanus †Melursus theobaldi Ursus †U. deningeri †U. dolinensis †U. etruscus †U. ingressus †U. kanivetz †U. kudarensis
Huracan_(mammal)
Miguelón or Skull 5. The site also contains thousands of bones from Ursus deningeri, ancestor of the cave bear, among those from other smaller carnivores
Paleolithic_Iberia
Extinct species of bear
†Aurorarctos †Protarctos †Helarctos sinomalayanus †Melursus theobaldi Ursus †U. deningeri †U. dolinensis †U. etruscus †U. ingressus †U. kanivetz †U. kudarensis
Ailuropoda_microta
Extinct species of bear
†Aurorarctos †Protarctos †Helarctos sinomalayanus †Melursus theobaldi Ursus †U. deningeri †U. dolinensis †U. etruscus †U. ingressus †U. kanivetz †U. kudarensis
Ailuropoda_baconi
Extinct genus of bears
†Aurorarctos †Protarctos †Helarctos sinomalayanus †Melursus theobaldi Ursus †U. deningeri †U. dolinensis †U. etruscus †U. ingressus †U. kanivetz †U. kudarensis
Miomaci
Extinct tribe of bears
†Aurorarctos †Protarctos †Helarctos sinomalayanus †Melursus theobaldi Ursus †U. deningeri †U. dolinensis †U. etruscus †U. ingressus †U. kanivetz †U. kudarensis
Ursavini
Extinct subfamily of bears
†Aurorarctos †Protarctos †Helarctos sinomalayanus †Melursus theobaldi Ursus †U. deningeri †U. dolinensis †U. etruscus †U. ingressus †U. kanivetz †U. kudarensis
Hemicyoninae
Homo erectus fossil
red fox (Vulpes vulpes), the wolf Canis mosbachensis, and the bear Ursus deningeri. Bears and humans possibly occupied the cave during different seasons
Tautavel_Man
Extinct species of carnivore
Ursus rossicus is an extinct species of bear that lived in the steppe regions of northern Eurasia and Siberia during the Pleistocene. It is part of the
Ursus_rossicus
Tribe of bears
†Aurorarctos †Protarctos †Helarctos sinomalayanus †Melursus theobaldi Ursus †U. deningeri †U. dolinensis †U. etruscus †U. ingressus †U. kanivetz †U. kudarensis
Agriotheriini
Extinct genus of bears
†Aurorarctos †Protarctos †Helarctos sinomalayanus †Melursus theobaldi Ursus †U. deningeri †U. dolinensis †U. etruscus †U. ingressus †U. kanivetz †U. kudarensis
Zaragocyon
Extinct genus of bears
†Aurorarctos †Protarctos †Helarctos sinomalayanus †Melursus theobaldi Ursus †U. deningeri †U. dolinensis †U. etruscus †U. ingressus †U. kanivetz †U. kudarensis
Kretzoiarctos
Extinct genus of bears
†Aurorarctos †Protarctos †Helarctos sinomalayanus †Melursus theobaldi Ursus †U. deningeri †U. dolinensis †U. etruscus †U. ingressus †U. kanivetz †U. kudarensis
Phoberogale
Cave in Turkey
of cave bears (a late form of Ursus deningeri) along with lower numbers of brown bears (a relatively early form of Ursus arctos). The cave bear and other
Yarımburgaz_Cave
Cave and archaeological site in the United Kingdom
den by cave bears (on the transition between the archaic Ursus deningeri and the later Ursus spelaeus) resulting in a considerable number of their remains
Kents_Cavern
Villalba de Alvarado et al. (2024) describe new fossil material of Ursus deningeri from the Salbatore II cave site (Spain), providing new information
2024_in_paleomammalogy
Prehistoric cave in Montbron, Charente, France
remains were also found. Layer 13 contains archaic fauna: Ursus deningeri (ancestor of Ursus spelaeus), Crocuta crocuta intermedia, and steppe rhinoceros
Montgaudier_Cave
specimen, and was also contaminated with DNA from an ancient cave bear (Ursus deningeri) present in the cave. The mtDNA analysis found a surprising link between
Molecular_paleontology
Extinct genus of bears
but have since been reassigned & synonymized with the Asian black bear (Ursus thibetanus). Various dental characteristics link Plionarctos with other
Plionarctos
Extinct genus of bears
†Aurorarctos †Protarctos †Helarctos sinomalayanus †Melursus theobaldi Ursus †U. deningeri †U. dolinensis †U. etruscus †U. ingressus †U. kanivetz †U. kudarensis
Dinocyon
Cave complex and archaeological site in the Czech Republic
steppe bison, but a few from aurochs), bears (primarily Ursus deningeri, but a few from Ursus spelaeus), reindeer, horses and wolves. More than 100 human
Mladeč_Caves
Extinct species of bear
floridanus is not considered a close relative of the Eurasian cave bear (Ursus spelaeus), which belonged to a different genus and subfamily. T. floridanus
Tremarctos_floridanus
Bocherens (2019). A study on the cranial and mandibular morphology of Ursus deningeri compared to other bear species, and on its implications for inferring
2019_in_paleomammalogy
Stone quarry in Thuringia, Germany
deer (Megaloceros sp.) Roe deer (Capreolus suessenbornensis) Bear (Ursus deningeri-spelaeus) Lion (Panthera spelaea) European wildcat (Felis silvestris)
Bilzingsleben (Paleolithic site)
Bilzingsleben_(Paleolithic_site)
Extinct genus of bears
†Aurorarctos †Protarctos †Helarctos sinomalayanus †Melursus theobaldi Ursus †U. deningeri †U. dolinensis †U. etruscus †U. ingressus †U. kanivetz †U. kudarensis
Cephalogale
Cave and archaeological site in southern France
and rhino (Dicerorhinus hemitoechus) and remains of Deninger bears (Ursus deningeri). Known since the middle of the nineteenth century for the remains
Arago_Cave
Period in the geologic temperature record
Long-eared bat ? Myotis mystacinus Whiskered bat ? Myotis bechsteini Bechstein's bat ? Scuirus sp. Squirrel ? Canis lupus Wolf ? Ursus deningeri † Bear ?
Marine_Isotope_Stage_13
Extinct genus of bears
Arctodus and Arctotherium, and an extinct subspecies of the modern polar bear Ursus maritimus tyrannus, Agriotherium was among the largest known terrestrial
Agriotherium
Extinct genus of bears
†Aurorarctos †Protarctos †Helarctos sinomalayanus †Melursus theobaldi Ursus †U. deningeri †U. dolinensis †U. etruscus †U. ingressus †U. kanivetz †U. kudarensis
Phoberocyon
Extinct genus of bears
†Aurorarctos †Protarctos †Helarctos sinomalayanus †Melursus theobaldi Ursus †U. deningeri †U. dolinensis †U. etruscus †U. ingressus †U. kanivetz †U. kudarensis
Plithocyon
Extinct genus of bears
†Aurorarctos †Protarctos †Helarctos sinomalayanus †Melursus theobaldi Ursus †U. deningeri †U. dolinensis †U. etruscus †U. ingressus †U. kanivetz †U. kudarensis
Ballusia
River in Spain
species in La Lucia cave (Lamasón, Cantabria, N Spain): Ursus deningeri von Reichenau and Ursus spelaeus Rosenmüller-Heinroth" (PDF). MUNIBE (Antropologia-Arkeologia):
Latarmá_River
Extinct genus of bears
†Aurorarctos †Protarctos †Helarctos sinomalayanus †Melursus theobaldi Ursus †U. deningeri †U. dolinensis †U. etruscus †U. ingressus †U. kanivetz †U. kudarensis
Hemicyon
Extinct genus of bears
†Aurorarctos †Protarctos †Helarctos sinomalayanus †Melursus theobaldi Ursus †U. deningeri †U. dolinensis †U. etruscus †U. ingressus †U. kanivetz †U. kudarensis
Ursavus
Extinct genus of bears
†Aurorarctos †Protarctos †Helarctos sinomalayanus †Melursus theobaldi Ursus †U. deningeri †U. dolinensis †U. etruscus †U. ingressus †U. kanivetz †U. kudarensis
Indarctos
Extinct genus of mammals
†Aurorarctos †Protarctos †Helarctos sinomalayanus †Melursus theobaldi Ursus †U. deningeri †U. dolinensis †U. etruscus †U. ingressus †U. kanivetz †U. kudarensis
Amphicynodon
Jiangzuo et al. (2018), who unambiguously confirm the presence of Ursus deningeri in Loc. 1 of Zhoukoudian. A study on the bone histology of cave bear
2018_in_paleomammalogy
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URSUS DENINGERI
URSUS DENINGERI
URSUS DENINGERI
URSUS DENINGERI
URSUS DENINGERI
URSUS DENINGERI
URSUS DENINGERI
URSUS DENINGERI
n.
Any species of the genus Ursus, and of the closely allied genera. Bears are plantigrade Carnivora, but they live largely on fruit and insects.
n.
The urus.
n.
The urus.
n.
The bearded seal.
n.
The Hebrew name of a horned wild animal, probably the Urus.
n.
A genus of Carnivora including the common bears.
n.
The European bison (Bison bonasus, / Europaeus), once widely distributed, but now nearly extinct, except where protected in the Lithuanian forests, and perhaps in the Caucasus. It is distinct from the Urus of Caesar, with which it has often been confused.
n.
A very large, powerful, and savage extinct bovine animal (Bos urus / primigenius) anciently abundant in Europe. It appears to have still existed in the time of Julius Caesar. It had very large horns, and was hardly capable of domestication. Called also, ur, ure, and tur.
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