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Frazione of Italy
45°35′43″N 11°12′36″E / 45.59528°N 11.21000°E / 45.59528; 11.21000 Monte Bolca is an Early Eocene-aged geologic site located near Verona, Italy. A Konservat-Lagerstätte
Monte_Bolca
Village in Vestenanova, Verona, Italy
Bolca is a village in the Veneto, on the southern margin of the Italian Alps. It is a frazione of the comune of Vestenanova, in the province of Verona
Bolca
The Eocene-aged Monte Bolca fossil site near Verona, Italy was one of the first lagerstatte to be discovered to science, and still has one of the most
Paleobiota_of_Monte_Bolca
Sedimentary deposit with well-preserved extraordinary fossils
Eocene Fur Formation, Green River Formation, Messel Formation and Monte Bolca, the Miocene Foulden Maar and Ashfall Fossil Beds, the Pliocene Gray Fossil
Lagerstätte
Extinct genus of ray-finned fishes
extinct genus of billfish from the Eocene. It is only known from the Monte Bolca deposits in Italy, and was likely restricted to shallow, tropical waters
Blochius
Extinct genus of cartilaginous fishes
thomyorkei, known from fossil remains from the Early Eocene-aged Monte Bolca Lagerstätte of Italy. The specific epithet references the English musician
Dasyomyliobatis
Extinct genus of fishes
species Chiarachromis salazzarii is known only from the Early Eocene Monte Bolca lagerstätte of Northern Italy. Bellwood, David R.; Bannikov, Alexandre F
Chiarachromis
Extinct genus of fishes
paradoxum Agassiz, 1844 (type species) - Early Eocene of Italy (Monte Bolca) The type species, A. paradoxum, was initially identified in 1796 by Giovanni
Amphistium
Family of fishes
fish (Perciformes) from the Eocene of Bolca, Northern Italy" (PDF). Studi e ricerche sui giacimenti terziari di Bolca. XV (12): 15–22. Bannikov, A. F. (2000)
Lutjanidae
Family of ray-finned fishes
Mediterranean). An extinct relative, Callipteryx, is known from the Monte Bolca lagerstätte of the Lutetian epoch. Weevers are sometimes used as an ingredient
Weever
Species of fish
tetraodontid bony fish that lived from the Lutetian epoch of Eocene Monte Bolca. In life, it would have resembled a somewhat-flattened boxfish with five
Spinacanthus
Family of marine fish that can make powerful, self-propelled leaps out of water
The earliest fossil flyingfish are known from the Early Eocene of Monte Bolca, Italy. They include the genus Rhamphexocoetus Bannikov et al., 1985, which
Flying_fish
Order of ray-finned fishes
fossil specimens of Acropoma are known from the Early Eocene of Italy (Monte Bolca). All species of Acropomatiformes live in the sea, some also live in brackish
Acropomatiformes
Suborder of fishes
Eocene of Bolca in northern Italy, and a new genus for Psettopsis latellai Bannikov, 2005". Studi e ricerche sui giacimenti terziari di Bolca: p. 5-15
Flatfish
Extinct genus of fishes
Agassiz, 1834, A. filamentosus Agassiz, 1834), known from the famous Monte Bolca site in Italy. It is the only genus in the extinct family Acanthonemidae
Acanthonemus
Species of fish
Bolcabalistidae to contain both Eospinus and the genus Bolcabalistes from Monte Bolca as close relatives of both triggerfishes and boxfishes. The similar Moclaybalistes
Eospinus
Extinct family of fishes
tetraodontid bony fish that lived from the Lutetian epoch of Eocene Monte Bolca. In life, either genus would have resembled a somewhat-flattened boxfish
Spinacanthidae
Extinct genus of highly unusual stingray
known from three specimens, one nearly complete, from the famous Monte Bolca lagerstätte. The specimen was initially assigned as the holotype of Arechia
Lessiniabatis
Extinct genus of fishes
single species, C. speciosus (=C. recticaudus), known from the famous Monte Bolca site of Italy. It was initially thought to have been a relative of weeverfishes
Callipteryx
Fourth-level administrative divisions of Italy
View of Bolca, a frazione of the comune of Vestenanova, Veneto
Frazione
Suborder of fishes
Formation. A diversity of stingray fossils is known from the Eocene Monte Bolca formation from Italy, including the early stingaree Arechia, as well as
Stingray
Extinct genus of fishes
It contains a single species, D. vestenae, known from the famous Monte Bolca site of Italy. It is the only member of the family Ductoridae. It is the
Ductor
Extinct genus of prehistoric marine eel
two species: Bolcyrus bajai Blot, 1978 – Late Ypresian of Italy (Monte Bolca) Bolcyrus formosissimus (Eastman, 1905) – Late Ypresian of Italy and possibly
Bolcyrus
Extinct genus of fishes
Ypresian of Monte Bolca, Italy (=Amia indica Volta, 1796, Smerdis micracanthus Agassiz, 1833) †C. spinosum Agassiz, 1833 - Ypresian of Monte Bolca, Italy (not
Cyclopoma
Extinct genus of fishes
Bajaichthys is an extinct Ypresian (early Eocene) zeid from the Monte Bolca Lagerstätten of Italy. It contains a single species, B. elegans, and is the
Bajaichthys
Genus of sharks
cuvieri is known from extremely well-preserved full-body specimens from Monte Bolca, Italy. The oldest known species is G. glickmani from the Cenomanian of
Galeorhinus
Extinct species of fish
anglerfishes. It was described by Louis Agassiz in 1835 from the Monte Bolca locality. It became extinct during the middle Eocene (lowermost Lutetian)
Caruso_brachysomus
Genus of extinct requiem shark
bolcensis. It is known from multiple articulated individuals from the Bolca Konservat−Lagerstätte of the Ypresian of Italy. It was a shallow water species
Eogaleus
Family of fishes (anglerfish)
(named after the album Tarkus) is known from the Early Eocene-aged Monte Bolca fossil site of Italy. Ogcocephaloidei batfishes are dorsoventrally compressed
Ogcocephalidae
Extinct teleost fish genus
species of percomorph teleost fish from the Eocene (Late Ypresian-aged) Monte Bolca site in Italy. D. volans is the only species in the genus Dibango, known
Dibango_volans
Extinct genus of fishes
(=E. subvespertilio (de Blainville, 1818)), from the Ypresian-aged Monte Bolca site of Italy. As its name suggests are closely allied to the extant genus
Eoplatax
Family of fishes
anchovy is of the stem-engrauline Eoengraulis from the Early Eocene of Monte Bolca, Italy. Despite their abundance in the modern day and their tendency to
Anchovy
Family of fishes
highest diversity has been found in the northern Italian formation at Monte Bolca, formed from the sedimentation of the Tethys Ocean in the early Eocene (50-49
Frogfish
Extinct genus of fishes
tetraodontid bony fish that lived from the Lutetian epoch of Eocene Monte Bolca. In life, it would have resembled a compressed boxfish with five massive
Protobalistum
Family of octopuses
in Lebanon, and Bolcaoctopus from the Early Eocene (Ypresian) of Monte Bolca, Italy. The following is a maximum likelihood phylogenetic tree based on
Octopodidae
Genus of ray-finned fishes
Paleocene of Peru, resemble later species, such as M. rhombea of the Monte Bolca lagerstätte, and even the living species, M. maculata. Experts remain undecided
Mene
Extinct species of fish
It contains a single species, E. sorbinii from the Ypresian-aged Monte Bolca site in Italy. It had two pairs of long spines, one over each eye, and one
Eolactoria
Extinct genus of fishes
Eocene. It contains two species, both known from the Ypresian-aged Monte Bolca site of Italy. The following species are known: B. alessandroi Bannikov
Bradyurus
Extinct genus of fishes
volans is an extinct bony fish that lived during the Lutetian epoch of Monte Bolca. It combines features of both halfbeaks (i.e., its elongated lower jaw)
Rhamphexocoetus
Extinct species of fish
Palaeopomacentrus orphae is an extinct species of fish from the Ypresian epoch of Monte Bolca. It is the only known species of the genus Palaeopomacentrus. Bellwood,
Palaeopomacentrus
Extinct genus of snakes
an extinct species of marine palaeophiid snake from the Eocene of Monte Bolca, Italy. It had the highest vertebral count known among snakes, with 565
Archaeophis
Genus of fishes (fossil)
Eocene. It contains a single species, C. bicornis, from the famous Monte Bolca site in Italy. It is the only known member of the extinct scombriform family
Carangodes
Extinct genus of fishes
is known from a single species, A. longirostris, from the famous Monte Bolca site of Italy. It was a member of Centriscidae, making it a relative of
Aeoliscoides
Extinct genus of fishes
contains a single species, D. brevicauda, from the late Ypresian-aged Monte Bolca lagerstatten of Italy. It is thought to be closely related to the sympatric
Dalpiazella
Family of ray-finned fishes
(Early Eocene of Italy) Fossil wrasses date to the Early Eocene of Monte Bolca, Italy. Among these is Phyllopharyngodon, which can uniquely be placed in
Wrasse
Family of pufferfish
Middle Eocene Europe, with fossils found in Monte Bolca and the Caucasus Mountains. The Monte Bolca species, E. pygmaeus, coexisted with several other
Tetraodontidae
Extinct species of fish
the Early Eocene. It contains a single species, C. pinnatiformis of Monte Bolca, Italy. It and Vomeropsis are the only known members of the extinct subfamily
Ceratoichthys
Order of fishes
Paranguilla, an Eocene eel from Monte Bolca
Eel
Extinct genus of fishes
definitive species, E. minimus (Agassiz, 1839) from the Early Eocene-aged Monte Bolca site of Italy. This species was initially erroneously described by Volta
Eobothus
Extinct genus of fishes
early Eocene. It contains a single species, B. leptacanthus, from the Monte Bolca lagerstatten of Italy. It was a member of the Holocentridae, making it related
Berybolcensis
Family of cartilaginous fishes
almost complete and articulated material from the Konservat-Lagerstätten of Bolca in Italy suggested that †"Rhinobatos" dezignii and †"Rhinobatos" primaevus
Guitarfish
Family of fishes
two fossil genera are also known, both from the Early Eocene-aged Monte Bolca site of Italy: †Histionotophorus Eastman, 1904 †Orrichthys Carnevale & Pietsch
Handfish
Order of fishes
Carlomonnius Bannikov & Carnevale, 2016 from the Early Eocene-aged Monte Bolca site of Italy. This genus was previously considered an indeterminate gobioid
Gobioidei
Extinct genus of fishes
early Eocene of Italy (Monte Bolca lagerstatten) †E. tavernei Tyler & Bannikov, 2012 - early Eocene of Italy (Monte Bolca lagerstatten) As the name of
Eotetraodon
species, Chalcophora eocenica sp. nov. from the lower Eocene of Pesciara (Bolca), northern Italy and revision of Perotis laevigata Heer, 1847 sensu Massalongo
2026_in_paleoentomology
Extinct genus of ray-finned fishes
family Siganidae. It is known from the Eocene lagerstätte "Pesciara" in Bolca, Italy. "†Acanthopygaeus Leriche 1906". Paleobiology Database. Fossilworks
Acanthopygaeus
Genus of fishes
species "Dules" temnopterus Agassiz, 1836 from the Early Eocene of Monte Bolca, Italy, was formerly placed in this genus. However, more recent analyses
Dules
Species of fish
Lichia veronensis Bannikov, 1990, is known from the Early Eocene-aged Monte Bolca site of Italy, suggesting that the genus has inhabited its present range
Leerfish
Family of fishes with caudal spines
particularly large diversity of fossil surgeonfish is known from the Monte Bolca Lagerstätte of Italy. These represent some of the earliest representatives
Acanthuridae
Extinct species of fish
barracuda known from the Eocene. It is known from the late Ypresian-aged Monte Bolca site of Italy. It is one of the earliest and best-known fossil barracudas
Sphyraena_bolcensis
Family of fishes
are known as fossils from the Early Eocene-aged lagerstatten of Monte Bolca, Eocene, in Italy. The genus Schuettea closely resembles members of Monodactylus
Monodactylidae
Family of cartilaginous fishes
Campanian/Maastrichtian of Nardo, Italy, and Eoplatyrhina from the Ypresian of Monte Bolca, Italy. Genus Platyrhina J. P. Müller & Henle, 1838 Platyrhina hyugaensis
Platyrhinidae
Extinct genus of fishes
common form of fossil. Valid species of Pycnodus is only known from Monte Bolca Lagerstätte in Italy, however fossils that were assigned to Pycnodus have
Pycnodus
Order of fishes
containing only Rhamphognathus) are known from the Early Eocene of Monte Bolca, Italy. Uniquely, these large-sized atheriniforms appear to have adapted
Atheriniformes
Extinct genus of fishes
Eocene epoch. It contains a single species, B. brachycephala from the Monte Bolca site of Italy. Its exact taxonomic affinities within the Anguilliformes
Bolcanguilla
"†Neilpeartia ceratoi, gen. et sp. nov., a new frogfish from the Eocene of Bolca, Italy". Journal of Vertebrate Paleontology. 40 (2) e1778711. Bibcode:2020JVPal
List of organisms named after famous people (born 1950–1974)
List_of_organisms_named_after_famous_people_(born_1950–1974)
Extinct genus of ray-finned fishes
Ypresian epoch in what is now Monte Bolca, northern Italy. It is one of two known fossil Moorish idols from Monte Bolca alongside Angiolinia. It differs
Eozanclus
Extinct genus of fishes
Eocene epoch. It contains a single species, G. homopterus from the Monte Bolca site of Italy. Its exact taxonomic affinities within the Anguilliformes
Gazolapodus
Family of ray-finned fishes
fish from the Eocene of Bolca in northern Italy (Periformes, Trichiuroidea)." Studi e Ricerche sui giacimenti Terziari di Bolca, XII Miscellanea Paleontologica
Gempylidae
Extinct species of fish
Monte Bolca, about 30 kilometres north-east of Verona, Italy. Restoration comparing †Mene rhombea and †Mene oblonga Mene rhombea, from Monte Bolca, Verona
Mene_rhombea
Canadian drummer and lyricist (1952–2020)
"†Neilpeartia ceratoi, gen. et sp. nov., a new frogfish from the Eocene of Bolca, Italy". Journal of Vertebrate Paleontology. 40 (2) e1778711. Bibcode:2020JVPal
Neil_Peart
Family of fishes
(2012). "†Caruso, a new genus of anglerfishes from the Eocene of Monte Bolca, Italy, with a comparative osteology and phylogeny of the teleost family
Goosefish
contains a single species, M. gigas, known from the Early Eocene of Monte Bolca, Italy. Its genus name, Monopteros meaning "one wing", comes from its large
Monopteros_(fish)
Genus of fishes
Bannikov, Alexandre F.; Marramà, G.; Tyler, James C.; Zorzin., R. (2014). "The Bolca Fossil-Lagerstätte: A window into the Eocene World. 5. The Pesciara- Monte
Trumpetfish
Family of ray-finned fishes
†Trollichthys Marramà & Carnevale, 2015 from the Early Eocene of Monte Bolca, Italy, which was initially considered a round herring, has been identified
Spratelloididae
Extinct species of fish
an extinct species of ray-finned fish that first appeared in the Monte Bolca Lagerstatten during the Lutetian epoch of the Eocene. For a menid, it has
Mene_oblonga
Extinct species of shark
an extinct species of houndshark known from the Early Eocene-aged Monte Bolca site of Italy. It was a close relative of the modern tope or school shark
Galeorhinus_cuvieri
Family of boxfishes
†Eolactoria Tyler, 1975 †Eolactoria sorbinii Tyler 1976 (Lutetian of Monte Bolca, Eocene Italy) †Oligolactoria Tyler & Gregorova, 1991 †Oligolactoria bubiki
Ostraciidae
Extinct genus of fishes
nearly complete specimen recovered from the Monte Postale site of the Monte Bolca locality. It was first named by Alexandre F. Bannikov and Giorgio Carnevale
Pietschellus
Extinct prehistoric eel that lived during the Lutetian epoch of the Eocene
that lived during the Lutetian epoch of the Eocene, in what is now Monte Bolca. Some of the better preserved fossils show that the living animals had complex
Paranguilla
Species of ray-finned fish
FROM BOLCA, NORTHERN ITALY, PREVIOUSLY CLASSIFIED WITH THE APOGONIDAE AND ENOPLOSIDAE (PERCIFORMES)". Studi e ricerche sui giacimenti terziari di Bolca. 12:
Old_wife
Family of cartilaginous fishes
Titanonarke is known from complete skeletons from the Early Eocene-aged Monte Bolca site of Italy. At nearly 1 metre (3.3 ft) in length, it is the largest narcinid
Narcinidae
Extinct genus of fishes
that lived in a coral reef during the Lutetian epoch of what is now Monte Bolca, Italy. Paleontology portal Fish portal Prehistoric fish List of prehistoric
Tylerichthys
Ray-finned fish with a natural luring appendage
earliest fossils of anglerfish are from the Eocene, excavated from the Monte Bolca formation of Italy, and these already show evidence of diversification into
Anglerfish
Extinct genus of fishes
Italy. Several exquisitely preserved fossils have been found from the Monte Bolca lagerstatten. Some juvenile specimens preserve the vertical striped coloration
Archaephippus
Extinct genus of ray-finned fishes
remains are primarily known from the late Ypresian-aged rocks of Monte Bolca, Italy. However, specimens have also been recovered from the earliest-Ypresian
Platinx
Extinct genus of fishes
prehistoric species of moonyfish that lived during the Lutetian epoch of Monte Bolca, Italy. It had large, rounded dorsal and anal fins, and was a comparatively
Psettopsis
Family of fishes
Caproidae (Acanthomorpha)". Studi e ricerche sui giacimenti terziari di Bolca. 11: 7–74. Maile, Alex J.; Smith, W. Leo; Davis, Matthew P. (2025-05-02)
Caproidae
Topics referred to by the same term
Rio, Florida, a community in the United States The "Pesciara" of Monte Bolca, a site famous for fish fossils near Verona, Italy Fishbowl (secure phone)
Fishbowl
Second epoch of the Paleogene Period
Calcareous nannoplankton were a prominent feature of Eocene marine ecosystems. Bolca in Italy List of fossil sites (with link directory) London Clay Messel pit
Eocene
Family of ray-finned fishes
Carnevale and James C. Tyler in 2024 based on three specimens found in Bolca, Italy. Carnevale & Tyler found that Zanclus cornutus and Angiolinia mirabilis
Zanclidae
Extinct genus of fishes
aracanid boxfish that lived during the Lutetian of middle Eocene Monte Bolca. Paleontology portal Fish portal Eolactoria Prehistoric fish List of prehistoric
Proaracana
Family of ray-finned fishes
PMID 25543675. Bannikov, Alexander F. (1984). "The New Subfamily of Carangid Fishes". Studi e ricerche sui giacimenti terziari di Bolca. IV: 319–321.
Carangidae
Genus of cartilaginous fishes
†Plesiozanobatus Marramà et al, 2020 is known from the Early Eocene-aged Monte Bolca site of Italy. Last; White; de Carvalho; Séret; Stehmann; Naylor, eds. (2016)
Panray
Extinct genus of fishes
sorbinii is an extinct sailfin moonfish from the Lutetian epoch of the Monte Bolca lagerstatten. Paleontology portal Carnevale, Giorgio (2018). "A sailfin
Veronavelifer
Chemical compound
Licoflavone C Xanthohumol, the corresponding prenylated chalcone Possemiers S, Bolca S, Grootaert C, Heyerick A, Decroos K, Dhooge W, De Keukeleire D, Rabot
Isoxanthohumol
Extinct genus of fishes
Monte Bolca Fossil of Sparnodus macrophthalmus from Monte Bolca Fossil of Sparnodus micracanthus Bannikov, A. 2006. Fishes from the Eocene of Bolca, northern
Sparnodus
Suborder of ray-finned fishes
indeterminate scorpaenoid is also known from the Early Eocene-aged Monte Bolca site in Italy. The suborder Scorpaenoidei is classified into families and
Scorpaenoidei
Family of fishes
tetraodontiform fishes from the Eocene of Monte Bolca, Italy, with comments on related Tertiary taxa". Studi e Ricerche sui Giacimenti Terziari di Bolca. 9: 47–119.
Aracanidae
Extinct genus of prehistoric bony fish
It contains a single species, E. leptoptera from the Ypresian-aged Monte Bolca site of Italy. It is thought to have been an early marine relative of the
Eoanguilla
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