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Archaic human species from Asia
The Denisovans or Denisova hominins (/dəˈnɪsəvə(nz)/ də-NIS-ə-və(nz)) are an extinct species or subspecies of archaic human that ranged across Asia during
Denisovan
Human hybridization during the Paleolithic
Interbreeding between archaic humans (such as Neanderthals and Denisovans) and anatomically modern humans (contemporaneous Homo sapiens) took place during
Interbreeding between archaic and modern humans
Interbreeding_between_archaic_and_modern_humans
Archaic human from China, 146,000 BP
belong to the same species as the Denisovans, and subsequent proteomics and mitochondrial DNA analyses confirmed its Denisovan affinities. The Harbin cranium
Harbin_cranium
Hominin fossil
000 year old fossil specimen belonging to a ~13-year-old Neanderthal-Denisovan hybrid girl. To date, she is the only first-generation hybrid hominin
Denny_(hybrid_hominin)
Cave and archaeological site in Russia
about 10,000 years earlier from South Africa from c. 61,000 years ago). Denisovans, Neanderthals and related hybrids may have inhabited the Denisova Cave
Denisova_Cave
Melanesian inhabitants of New Guinea
due to their Denisovan admixture. It is suggested that Papuans may have underwent a secondary admixture event with Altai-related Denisovan populations
Indigenous people of New Guinea
Indigenous_people_of_New_Guinea
Genetic study of ancient Neanderthal DNA
human females and Neanderthal males. Neanderthals also interbred with Denisovans in the Siberian Altai Mountains. Genetic studies on Neanderthal ancient
Neanderthal_genetics
One of the two categories of Indigenous Australians
with Altai-related Denisovan populations after they diverged from the ancestors of East Asians, who already mixed with Denisovans. This event was also
Aboriginal_Australians
Extinct species of human
humans, Neanderthals are most closely related to Denisovans based on nuclear DNA (nDNA) analyses. Denisovans are an enigmatic group of Late Pleistocene humans
Neanderthal
Evolutionary process
derive from Denisovans, indicating limited interbreeding in Southeast Asia. Alleles thought to have originated in Neanderthals and Denisovans have been
Human_evolution
Genus of hominins
remains poorly done. This has led to using common names ("Neanderthal" and "Denisovan"), even in scientific papers, to avoid trinomial names or the ambiguity
Homo
Archaic human from Luzon, Philippines
closely related to or directly a member of the Denisovan taxon, based on the unusually high percentage of Denisovan DNA found in Ayta Magbukon populations (southern
Homo_luzonensis
Hominin fossil
sequences in 2025 confirmed that the mandible represented that of a male Denisovan. The fossil was recovered sometime before 2008 by fishermen working in
Penghu_1
Spread of humans from Africa across the world
which lived around 500,000 years ago and was the likely ancestor of Denisovans and Neanderthals as well as modern humans. Early hominids had likely crossed
Early_human_migrations
Continent
Indian Ocean. Modern humans interbred with an archaic human species called Denisovans in Southeast Asia. Before the arrival of modern humans, Flores was occupied
Asia
Buddhist sanctuary and paleoanthrological site on the Tibetan Plateau in Gansu, China
way of palaeoproteomic analysis, is the first confirmed discovery of a Denisovan fossil outside of Denisova Cave. This fossil discovery shows that archaic
Baishiya_Karst_Cave
Hominin fossil
first confirmed discovery of a Denisovan fossil outside of Denisova Cave, and was the most complete confirmed Denisovan fossil before the identification
Xiahe_mandible
Extinct species of archaic human
as either the last common ancestor of modern humans, Neanderthals, and Denisovans; or as a completely separate lineage. H. heidelbergensis was described
Homo_heidelbergensis
Paintings, often prehistoric, on cave walls and ceilings
the oldest of such paintings were created not by Homo sapiens, but by Denisovans and Neanderthals, though this is contested. Discussion around prehistoric
Cave_painting
Chun-Hsiang; Cappellini, Enrico; Welker, Frido (2025-04-11). "A male Denisovan mandible from Pleistocene Taiwan". Science. 388 (6743): 176–180. Bibcode:2025Sci
List of human evolution fossils
List_of_human_evolution_fossils
Subregion of Oceania
humans, dubbed 'Denisovans', interbred with us BBC News online (2010-12-22) report (with video) on study that shows that Denisovans interbred with the
Melanesia
Extinct species of archaic human
heidelbergensis—the last common ancestor of modern humans, Neanderthals, and Denisovans. As such a widely distributed species both geographically and temporally
Homo_erectus
Country in Eastern Europe and North Asia
oldest Denisovan specimen lived 195–122,700 years ago. Fossils of Denny, an archaic human hybrid that was half Neanderthal and half Denisovan, and lived
Russia
Species of hominid in the genus Homo
non-sub-Saharan African populations, and Neanderthals and other hominins, such as Denisovans, may have contributed up to 6% of their genome to present-day non-sub-Saharan
Human
Ethnic group of the Philippines
into Philippine Negritos from Denisovans. The Ayta Magbukon in particular were found to possess the highest level of Denisovan ancestry in the world (between
Aeta_people
Archaeogenetic name for an ancestral genetic component
inherited their Denisovan ancestry from the same ancestral populations that ANE-related and Tianyuan-related populations inherited their Denisovan ancestry from
Ancient_North_Eurasian
Subregion of the Asian continent
Panninge individual branching off from the Near Oceanian clade after the Denisovan gene flow, although with the most supported topology indicating around
Southeast_Asia
Geographical region in the Pacific Ocean
in 2010, while no Eurasians or Africans displayed contributions of the Denisovan genes. The original inhabitants of the group of islands now named Melanesia
Oceania
Indigenous peoples from Mindanao in the Philippines
Southeast Asia. This is similar to the Sama Bajau people. The Manobo possess Denisovan admixture, much like the Mamanwa. Lumad Indigenous peoples of the Philippines
Manobo
Species of archaic human
which would encompass Xujiayao, Xuchang Man, Penghu 1, and possibly the Denisovans. Denisova Cave Baishiya Karst Cave Tam Ngu Hao 2 Cave Penghu 1 Xuchang
Homo_juluensis
Hominin fossil from China
people of Japan. A 2025 study states that the Tianyuan Man had the highest Denisovan ancestry among East Eurasians. Ancient and modern East Asians derive most
Tianyuan_man
Archaeological scientist
human dispersals and extinctions across Eurasia, including Neanderthals, Denisovans and modern Homo sapiens. Douka completed a B.Sc. in archaeological conservation
Katerina_Douka
Old Stone Age ''Homo sapiens''
Denisovans caused by a genetic bottleneck of the latter was dated at 744,000 years ago, combined with repeated early admixture events and Denisovans diverging
Early_modern_human
Archipelagic country in Southeast Asia
displaced by later waves of migration. Some Philippine Negritos have a Denisovan admixture in their genome. Ethnic Filipinos generally belong to several
Philippines
Country in South Asia
Australia, Tasmania, and Papua New Guinea), picking up Neanderthal and Denisovan genes along the way; branches reached Europe somewhat later, and Americas
India
the Denisovans, discovered from mitochondrial and nuclear genomes recovered from bone found in a cave in Siberia. Analysis showed that the Denisovan specimen
Molecular_paleontology
Colonisation of the European continent
anatomically modern humans and the archaic sister species Neanderthals and Denisovans, has been described at Petralona Cave in Greece, Sima de los Huesos in
Human_dispersals_in_Europe
Hominid skull fossil from Yunxian, China
team of researchers to believe the fossils are ancestral relatives of Denisovans. The first two skulls bear similarities to Dali Man, but are significantly
Yunxian_Man
sapiens Peștera cu Oase Lagar Velho 1 Contemporary species Homo erectus Denisovan Homo floresiensis Homo naledi Homo sapiens Research Excavation sites Neanderthal
Neanderthal_extinction
Mountain range in Central–East Asia
period of the Last Interglacial. The Altai mountains were home to the Denisovan branch of hominids who were contemporaries of Neanderthals and of Homo
Altai_Mountains
Protein found in humans
adaptation in humans. One such allele, which has been inherited from Denisovan archaic hominins, is known to confer increased athletic performance in
EPAS1
People of Andaman archipelago
genetically roughly equidistant to Denisovans and Neanderthals. Other studies state that Andaman Islanders have Denisovan ancestry but it's lower than what's
Andamanese_peoples
Region in central Asia
2010 discovery of 50,000-year-old fossils of a new kind of human, the Denisovans. Since then, Neanderthal bones, and tools crafted by Homo sapiens have
Altai-Sayan_region
Research institute based in Leipzig, Germany
recently published Neanderthal genome. A new population was described, Denisovans, which contributed 4–6% of its genome to modern Melanesians, similar to
Max Planck Institute for Evolutionary Anthropology
Max_Planck_Institute_for_Evolutionary_Anthropology
Archaic human species from 2.4 to 1.65 mya
ergaster) (Au. sediba) H. heidelbergensis Homo sapiens Neanderthals Denisovans ← Earlier apes ← Gorilla split ← Chimpanzee split ← Earliest bipedal
Homo_habilis
Hominin fossil
Asian contemporaries represent these enigmatic "Denisovans", but this is impossible to confirm as Denisovans are only identifiable from DNA instead of any
Dali_Man
Island and region in Indonesia
Southeast Asian populations indicates the widespread dispersal of the Denisovans (a species currently recognisable only by their genetic signature) across
Java
Fossil site in San Diego County, California
ergaster) (Au. sediba) H. heidelbergensis Homo sapiens Neanderthals Denisovans ← Earlier apes ← Gorilla split ← Chimpanzee split ← Earliest bipedal
Cerutti_Mastodon_site
Front of head
cold climate via fat distribution – introgressed from ancient humans – Denisovans – into the modern humans Native Americans. Another study found look-alike
Face
Hominin fossil
H. ergaster (?) H. floresiensis Penghu 1 H. heidelbergensis H. longi (Denisovans) H. luzonensis H. neanderthalensis H. rhodesiensis (?) Modern humans Homo
Eve's_footprint
Aboriginal Australians and Torres Strait Islander peoples
Papuan and Mamanwa peoples carry some of the alleles associated with the Denisovan peoples of Asia, (not found amongst populations in mainland Asia) suggesting
Indigenous_Australians
1955 novel by William Golding
sapiens Peștera cu Oase Lagar Velho 1 Contemporary species Homo erectus Denisovan Homo floresiensis Homo naledi Homo sapiens Research Excavation sites Neanderthal
The Inheritors (Golding novel)
The_Inheritors_(Golding_novel)
Terrestrial locomotion using two limbs
ergaster) (Au. sediba) H. heidelbergensis Homo sapiens Neanderthals Denisovans ← Earlier apes ← Gorilla split ← Chimpanzee split ← Earliest bipedal
Bipedalism
Paleogeneticist
identified that a teenage girl born 90,000 years ago had both Neanderthal and Denisovan parents. She was selected as one of Nature's 10 in 2018. Slon completed
Viviane_Slon
Offspring of cross-species reproduction
and interbred with Neanderthals, Denisovans, and at least one other extinct Homo species. Thus, Neanderthal and Denisovan DNA has been incorporated into
Hybrid_(biology)
Speakers of Austronesian languages
broad groups based on Denisovan admixture. Philippine Negritos, Papuans, Melanesians, and Indigenous Australians display Denisovan admixture, while Malaysian
Austronesian_peoples
Overview of genetic research on Pleistocene epoch archaic and early modern humans"
and Denisovans (Homo denisova) diverged from a common ancestral population between c. 500-700 thousand years ago (kya). Neanderthals and Denisovans subsequently
Genetic history of Pleistocene humans
Genetic_history_of_Pleistocene_humans
American geneticist (born 1974)
Europe, and on archaic admixture between modern humans, neanderthals, and denisovans. He is a professor in the Department of Genetics at Harvard Medical School
David_Reich_(geneticist)
Extinct hominid from Miocene Africa
ergaster) (Au. sediba) H. heidelbergensis Homo sapiens Neanderthals Denisovans ← Earlier apes ← Gorilla split ← Chimpanzee split ← Earliest bipedal
Sahelanthropus
Prehistoric inhabitants in Japan
inherited their Denisovan ancestry from the same ancestral populations that ANE-related and Tianyuan-related populations inherited their Denisovan ancestry from
Jōmon_people
Archaeological period, part of Stone Age
ergaster) (Au. sediba) H. heidelbergensis Homo sapiens Neanderthals Denisovans ← Earlier apes ← Gorilla split ← Chimpanzee split ← Earliest bipedal
Middle_Paleolithic
French paleoanthropologist
to the Denisovan lineage, despite the absence of recoverable DNA. This discovery significantly expanded the known geographic range of Denisovans and demonstrated
Jean-Jacques_Hublin
National Historic Site of Tanzania
ergaster) (Au. sediba) H. heidelbergensis Homo sapiens Neanderthals Denisovans ← Earlier apes ← Gorilla split ← Chimpanzee split ← Earliest bipedal
Olduvai_Gorge
Transfer of genetic material from one species to another
There is strong evidence for the introgression of Neanderthal genes and Denisovan genes into parts of the modern human gene pool, and Homo sapiens as a
Introgression
Upper Paleolithic archaeological site in Russia
into the Upper Paleolithic Sunghir population, consistent with traces of Denisovan ancestry. This type of ancestry is absent from the Kostenki14 specimen
Sungir
Proposal that humans evolved certain features due to filling a semi-aquatic niche
H. ergaster (?) H. floresiensis Penghu 1 H. heidelbergensis H. longi (Denisovans) H. luzonensis H. neanderthalensis H. rhodesiensis (?) Modern humans Homo
Aquatic_ape_hypothesis
Caves in France containing Paleolithic paintings
H. ergaster (?) H. floresiensis Penghu 1 H. heidelbergensis H. longi (Denisovans) H. luzonensis H. neanderthalensis H. rhodesiensis (?) Modern humans Homo
Lascaux
Earliest anatomically modern humans in Europe and West Asia
Paleolithic Sunghir (34kya) population is consistent with a low amount of Denisovan ancestry among them, yet absent from the Kostenki14 specimen, which is
Cro-Magnon
Theory of early hominid migration
affinity within Neanderthals' mitochondrial genomes to Homo sapiens than Denisovans. An interpretation, with as of yet limited recognition, of a skull fragment
Recent African origin of modern humans
Recent_African_origin_of_modern_humans
Superfamily of primates
ergaster) (Au. sediba) H. heidelbergensis Homo sapiens Neanderthals Denisovans ← Earlier apes ← Gorilla split ← Chimpanzee split ← Earliest bipedal
Ape
high frequency of an EPAS1 haplotype acquired through introgression with Denisovans, which assists with adaptation to low oxygen levels at high elevation
Genetic history of East Asians
Genetic_history_of_East_Asians
Extinct population of archaic humans
mandible (lower jaw) Levallois flake found at Nesher Ramla Biology portal Denisovan – Archaic human species from Asia Homo longi – Archaic human species from
Nesher_Ramla_Homo
Biological process
used to reconstruct the full methylation maps of the Neanderthal and the Denisovan. In September 2019, researchers published a novel method to infer morphological
DNA_methylation
Japanese prehistorical period
M. (October 2025). "An early East Asian lineage with unexpectedly low Denisovan ancestry". Current Biology. 35 (20): 4898–4908.e4. doi:10.1016/j.cub.2025
Jōmon_period
Organisms capable of living at high altitudes
Tibetan population has been linked to Denisovan-related populations. The Tibetan haplotype is more similar to the Denisovan haplotype than any modern human
Organisms_at_high_altitude
Hominin fossil
Related species Homo erectus Homo antecessor Homo cepranensis Homo neanderthalensis Homo sp. Altai (Denisovans) Homo sapiens Evolutionary biology portal
Saldanha_man
Objects worn to cover the body
needle at least 50,000 years old from Denisova Cave in Siberia made by Denisovans. Dyed flax fibers that date back to 34,000 BC and could have been used
Clothing
Cave with prehistoric paintings in Spain
H. ergaster (?) H. floresiensis Penghu 1 H. heidelbergensis H. longi (Denisovans) H. luzonensis H. neanderthalensis H. rhodesiensis (?) Modern humans Homo
Cave_of_Altamira
Ethnic groups of Asia
from ancient Guangxi populations and Han Chinese and tenuously have no Denisovan introgression. These findings suggest that Hlai and Ami/Atayal form a
Kra–Dai-speaking_peoples
Winter et al., it is also present in the Vindija Neandertal and Altai Denisovan sequences. Hence the dating must be older than 700 00 kya and possibly
KRT41P
Archaeological site
additional genetic affinity for Denisovans that is not matched by modern Native Americans; this additional Denisovan affinity is likely due to sampling
Upward_Sun_River_site
mutant EPAS1 gene could have been inherited from archaic hominins, the Denisovans. EPAS1 and EGLN1 are believed to be important genes for unique adaptive
High-altitude adaptation in humans
High-altitude_adaptation_in_humans
Geographical region of Russia comprising North Asia
Siberia around 40,000 years ago: H. sapiens, H. neanderthalensis, and the Denisovans. In 2010, DNA evidence identified the last as a separate species. Late
Siberia
People of the Philippines
divergent from each other. Philippine Negritos show a high degree of Denisovan Admixture, similar to Papuans and Indigenous Australians, in contrast
Filipinos
Prehistoric humans from 12,500 BCE in southwest China
represent a late survival of an archaic human species, hybridization between Denisovan hominin and modern humans, or alternatively just "an unfortunate overinterpretation
Red_Deer_Cave_people
will replace several smaller types of nuclear bombs in its arsenal. 2008 Denisovans The third discovered kind of human. NS 50 Let Pobedy, the world's largest
Timeline of Russian innovation
Timeline_of_Russian_innovation
ergaster) (Au. sediba) H. heidelbergensis Homo sapiens Neanderthals Denisovans ← Earlier apes ← Gorilla split ← Chimpanzee split ← Earliest bipedal
Neanderthals_in_Gibraltar
Pre-Neanderthal population in Spain
that the individual shares a closer common ancestor with Central Asian Denisovans (the sister group of Neanderthals) than with Neanderthals. In 2016, however
Sima_de_los_Huesos_hominins
Effort to sequence the Neanderthal genome
with modern humans multiple times, and that Neanderthals interbred with Denisovans only once, as evidenced in the genome of modern-day Melanesians. In 2006
Neanderthal_genome_project
Topics referred to by the same term
1997 video game for the Sega Genesis Woman X, a name given to the first Denisovan discovered This disambiguation page lists articles associated with the
X_woman
Prehistoric period, first part of the Stone Age
ergaster) (Au. sediba) H. heidelbergensis Homo sapiens Neanderthals Denisovans ← Earlier apes ← Gorilla split ← Chimpanzee split ← Earliest bipedal
Paleolithic
Ethnic group of the Philippines
peopling of the Philippines. Furthermore, they display high percentages of Denisovan gene flow. Legends, such as those involving the Ten Bornean Datus and
Ati_people
Bag in Khongor, Darkhan-Uul, Mongolia
Matthias; Kelso, Janet; Peter, Benjamin M.; Pääbo, Svante (30 October 2020). "Denisovan ancestry and population history of early East Asians". Science. 370 (6516):
Salkhit
Mythical creature
2024 in the Adirondack Mountains and DNA testing showed it to be 58% Denisovan/Neanderthal and 41% modern human. Stuart invited scientists to examine
Bigfoot
Paleoanthropological site in northeastern China
; Welker, F.; Shen, C.-C.; et al. (2019). "A late Middle Pleistocene Denisovan mandible from the Tibetan Plateau" (PDF). Nature. 569 (7756): 409–412
Jinniushan
Transition of human species to anthropologically modern behavior
evolutionary success of Homo sapiens worldwide relative to Neanderthals, Denisovans, and other archaic humans. There are many other hypotheses on the evolution
Behavioral_modernity
Upper Paleolithic culture of Europe
Matthias; Kelso, Janet; Peter, Benjamin M.; Pääbo, Svante (30 October 2020). "Denisovan ancestry and population history of early East Asians". Science. 370 (6516):
Aurignacian
Subspecies of Homo erectus (fossil) discovered on the island of Java in 1891
H. ergaster (?) H. floresiensis Penghu 1 H. heidelbergensis H. longi (Denisovans) H. luzonensis H. neanderthalensis H. rhodesiensis (?) Modern humans Homo
Java_Man
Period of human history before records
ergaster) (Au. sediba) H. heidelbergensis Homo sapiens Neanderthals Denisovans ← Earlier apes ← Gorilla split ← Chimpanzee split ← Earliest bipedal
Prehistory
Archaeological site in Nigeria
H. ergaster (?) H. floresiensis Penghu 1 H. heidelbergensis H. longi (Denisovans) H. luzonensis H. neanderthalensis H. rhodesiensis (?) Modern humans Homo
Iho_Eleru
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