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Age in the Middle Triassic
The Ladinian is a stage and age in the Middle Triassic series or epoch. It spans the time between 241.464 Ma and ~237 Ma (million years ago). The Ladinian
Ladinian
Italian author (1923–2006)
Alex Moroder (Ortisei, 13 May 1923 – Ortisei, 11 November 2006) was an Italian activist. He was the son of sculptor Ludwig Moroder and Adele Moroder. In
Alex_Moroder
First period of the Mesozoic Era
during the Early and Middle Triassic. Sea level rise accelerated in the Ladinian, culminating with a sea level up to 50 metres (164 ft) above present-day
Triassic
Stage of the Triassic
the Olenekian Age (part of the Lower Triassic Epoch) and precedes the Ladinian Age. The stage and its name were established by Austrian geologists Wilhelm
Anisian
Shang (2026). Zhang et al. (2026) determine the age and duration of the Ladinian Xingyi Fauna on the basis of the study of astrochronology and cyclostratigraphy
2026_in_reptile_paleontology
Extinct genus of beetles
Notocupes Temporal range: Ladinian–Turonian PreꞒ Ꞓ O S D C P T J K Pg N Various specimens of Notocupes from the Middle Jurassic of China and mid-Cretaceous
Notocupes
Third and final epoch of the Triassic Period
P Triassic J Lopingian ET Middle Late Early J Induan Olenekian Anisian Ladinian Carnian Norian Rhaetian ← Triassic–Jurassic extinction event ← Scleractinian
Late_Triassic
Extinct family of molluscs
Aplococeratidae Temporal range: Anisian–Ladinian PreꞒ Ꞓ O S D C P T J K Pg N Scientific classification Kingdom: Animalia Phylum: Mollusca Class: Cephalopoda
Aplococeratidae
Genus of fossil pseudosuchian
extinct genus of paracrocodylomorph known from the Middle–Late Triassic (Ladinian–Carnian ages) Chañares Formation of Argentina. The genus contains a single
Shakajlura
Genus of reptile
Carinthiasuchus is an extinct genus of poposauroid pseudosuchian from the Ladinian to Early Carnian Partnach Formation of Carinthia, Austria. Carinthiasuchus
Carinthiasuchus
Extinct genus of reptiles
Chañares Formation of Argentina, within strata belonging to the latest Ladinian stage of the late Middle Triassic, or the earliest Carnian of the Late
Luperosuchus
Extinct genus of reptiles
Qianichthyosaurus is an extinct genus of ichthyosaur from the Ladinian and Carnian stages of the Late Triassic epoch. Its fossils have been found in southeastern
Qianichthyosaurus
Lithostratigraphic unit in Europe
Keuper group and the Germanic Trias supergroup. It was deposited during the Ladinian stage of the Triassic period. It lies above the Upper Muschelkalk and below
Erfurt_Formation
Earth's most severe extinction event
measured by fossil evidence was far less rapid, showing up in the late Ladinian. Their adaptive radiation after the extinction event resulted in forms
Permian–Triassic extinction event
Permian–Triassic_extinction_event
Genus of pseudosuchian archosaurs
Parvosuchus Temporal range: Middle–Late Triassic (Ladinian–Carnian), ~237 Ma PreꞒ Ꞓ O S D C P T J K Pg N Holotype skull Scientific classification Kingdom:
Parvosuchus
Large region of volcanic rock in Russia
Smithian–Spathian, Olenekian–Anisian, Middle-Late Anisian and Anisian–Ladinian extinction events. Today, the area is covered by about 7 million km2 (3 million sq mi)
Siberian_Traps
Feature in geology
Rhaetian 201.4–208.5 Keuper Norian 208.5–227.0 Carnian 227.0–237.0 Middle Ladinian 237.0–242.0 Muschelkalk Anisian 242.0–247.2 Bunter or Buntsandstein Lower
Bunter_(geology)
Extinct genus of reptiles
28 November 2014. Diedrich, C. (2009). "The vertebrates of the Anisian/Ladinian boundary (Middle Triassic) from Bissendorf (NW Germany) and their contribution
Nothosaurus
Geologic formation in Tanzania
Formation (also known as the Manda Beds) is a Middle Triassic (Anisian? or Ladinian) or Late Triassic (earliest Carnian?) geologic formation in Tanzania. It
Manda_Formation
Extinct genus of lycopsid plants in the Order Isoetales
plant groups, with the youngest records of the genus being from the early Ladinian of North China, around 241.0-241.6 million years ago, with the genus likely
Pleuromeia
Extinct genus of amphibians
fossil record of Gerrothorax pulcherrimus extends 35 million years from the Ladinian stage of the Middle Triassic to the Rhaetian stage of the Late Triassic
Gerrothorax
Genus of molluscs (fossil)
Europe, during the Triassic, from the upper-most Anisian to the lower Ladinian age. This ammonoid has a ceratitic suture pattern on its shell (smooth
Ceratites
Extinct genus of reptiles
extinct genus of rhynchocephalian reptile known from the Middle Triassic (Ladinian) of Germany. It currently the most primitive known rhynchocephalian, and
Wirtembergia
Group of Triassic rocks in the United Kingdom
Anisian to Ladinian in age. Formerly known as the Preesall Salt (named from the Lancashire village of Preesall), this member is Anisian to Ladinian in age
Mercia_Mudstone_Group
Second epoch of the Triassic period
Late Triassic Epoch. The Middle Triassic is divided into the Anisian and Ladinian ages or stages. Formerly the middle series in the Triassic was also known
Middle_Triassic
Extinct genus of amphibians
(salamanders and close relatives). It is known from the Middle to Upper Triassic (Ladinian-Carnian) aged Madygen Formation in Kyrgyzstan. The type species is T. sixtelae
Triassurus
Extinct genus of fishes
Anisian of Yunnan, China. (type species) M. andreetti Tintori et al., 2007 – Ladinian of Lombardy, Italy. M. faccii (Gortani, 1907) – Carnian of Friuli-Venezia
Marcopoloichthys
Extinct group of marine reptiles
(2020-11-15). "Footprints of marine reptiles from the Middle Triassic (Anisian-Ladinian) Guanling Formation of Guizhou Province, southwestern China: The earliest
Thalattosauria
First age of the Late Triassic epoch
from 237 to 227.3 million years ago (Ma). The Carnian is preceded by the Ladinian and is followed by the Norian. Its boundaries are not characterized by
Carnian
Extinct genus of pseudosuchians
Etjosuchus recurvidens, which is based on a partial skeleton from the Ladinian to Carnian-aged Omingonde Formation. The holotype of Etjosuchus, GSN F382
Etjosuchus
Geological formation in Italy and Switzerland
black shale, is famous for its preservation of Middle Triassic (Anisian–Ladinian) marine life including fish and aquatic reptiles. It is exposed in the
Besano_Formation
Clade of reptiles
Pseudosuchians appeared during the late Olenekian (early Triassic); by the Ladinian (late Middle Triassic) they dominated the terrestrial carnivore niches
Pseudosuchia
Second age of the late Triassic
; 2005: The Global boundary Stratotype Section and Point (GSSP) of the Ladinian Stage (Middle Triassic) at Bagolino (Southern Alps, Northern Italy) and
Norian
Extinct genus of reptiles
archosaur, belonging to the clade Aphanosauria, from the Middle Triassic (late Ladinian age) Lower Santa Maria Formation in Paleorrota Geopark, Brazil. Friedrich
Spondylosoma
Extinct genus of reptiles
Kudnu Temporal range: Middle Triassic (Ladinian) ~239 Ma PreꞒ Ꞓ O S D C P T J K Pg N Holotype skull QM F9181 (top) and referred skull QM F9182 (bottom)
Kudnu
Genus of molluscs (fossil)
Attila; Budai, Tamás; Bercsényi, Miklós; Pintér, Zsolt (2025-12-23). "Ladinian ammonoids from the Balaton Highland (Hungary) : II. Veszprémfajsz". Földtani
Argolites
Extinct genus of reptiles
what is now Queensland, Australia during the Middle to lowermost Late (Ladinian–Carnian) Triassic period (236–233 Mya). The genus is made up of a single
Eomurruna
Extinct genus of reptiles
Limestone (Carnian, Late Triassic) of California, and also the Muschelkalk (Ladinian, Middle Triassic) of Germany. John Campbell Merriam (1902) described it
Californosaurus
Superfamily of beetles
Australia. The extinct family Obrieniidae, with species dating from the Ladinian stage of the Triassic through to tentatively the Oxfordian, have sometimes
Weevil
Gen. et sp. nov Valid Lacerda, Schultz & Bertoni-Machado Triassic (late Ladinian/early Carnian) Santa Maria Formation Brazil A non-crocodylomorph loricatan
2015 in archosaur paleontology
2015_in_archosaur_paleontology
Extinct genus of dicynodonts
Shansiodon is a genus of dicynodont from Middle Triassic (Anisian and Ladinian) of China and South Africa (sp. indet.). v t e
Shansiodon
diversity deep marine ostracod assemblage from the Middle Triassic (early Ladinian) of the Northern Calcareous Alps (Austria)". Historical Biology: An International
2025 in arthropod paleontology
2025_in_arthropod_paleontology
Genus of basal avemetatarsalians
as the Makay Formation, which dates to the Middle/Late Triassic period (Ladinian–Carnian ages), though the exact numerical age is currently unknown. The
Mambachiton
Extinct genus of reptiles
from marine or coastal deposits of the Middle Triassic epoch (Anisian and Ladinian stages), with some exceptions. For example, a vertebra from Nova Scotia
Tanystropheus
Extinct genus of coelacanths
Ticinepomis Temporal range: Lower Ladinian, 240.91 Ma PreꞒ Ꞓ O S D C P T J K Pg N Fossil Reconstruction Scientific classification Kingdom: Animalia Phylum:
Ticinepomis
Type of limestone
Rhaetian 201.4–208.5 Keuper Norian 208.5–227.0 Carnian 227.0–237.0 Middle Ladinian 237.0–242.0 Muschelkalk Anisian 242.0–247.2 Bunter or Buntsandstein Lower
Buntsandstein
Geological formation in Austria and Italy
fossils dating back to the Middle Triassic period (late Anisian-middle Ladinian), or Illyrian to Longobardian in the regional stratigraphy. The formation
Schlern_Formation
Extinct genus of reptiles
genus of a diapsid reptile, known from the Middle Triassic (Anisian and Ladinian age) Besano Formation of northern Italy and Prosanto Formation of south-eastern
Eusaurosphargis
Extinct species of reptile
(distantly related to modern crocodilians) from the Middle Triassic (Anisian–Ladinian) of Switzerland and Italy. The genus name means "Ticino crocodile" due
Ticinosuchus
Extinct genus of reptiles
about 40 centimetres (16 in) long[citation needed] and lived during the Ladinian in the Middle Triassic, from about 242 to 235 million years ago. Cisneros
Candelaria_barbouri
Lithostratigraphic unit in Europe
Rhaetian 201.4–208.5 Keuper Norian 208.5–227.0 Carnian 227.0–237.0 Middle Ladinian 237.0–242.0 Muschelkalk Anisian 242.0–247.2 Bunter or Buntsandstein Lower
Muschelkalk
event. Lovegrove et al. (2025) describe a large silesaur femur from the Ladinian-Carnian Ntawere Formation (Zambia), and argue that the studied specimen
2025 in archosaur paleontology
2025_in_archosaur_paleontology
Extinct genus of reptiles
archosauromorph reptile known from the Middle Triassic (Late Anisian to Ladinian) of Europe and China. Macrocnemus is a member of the Tanystropheidae family
Macrocnemus
Species of liverwort
Klaus-Peter. "Naiadita lanceolata (Marchantiophyta) from the Middle Triassic (Ladinian) of Germany: a new reconstruction attempt and considerations on taphonomy"
Naiadita
Extinct genus of dicynodonts
Stahleckeria is an extinct genus of Middle Triassic (Ladinian) dicynodonts. It lived about 237 million years ago in what is now Brazil and Namibia. As
Stahleckeria
Extinct family of reptiles
family of basal marine reptiles known from the Middle Triassic (Anisian-Ladinian boundary) of southern Switzerland and northern Italy. The type species
Helveticosauridae
Extinct genus of reptiles
genus of diapsid marine reptile known from the Middle Triassic (Anisian-Ladinian boundary) of southern Switzerland and Italy. It contains a single species
Helveticosaurus
South Tyrolean journalist, writer and folklorist (1879-1966)
writer and self-taught folklorist known for collecting and publishing Ladinian legends. Karl Felix Wolff was born on 21 May 1879 in Karlstadt, Austria-Hungary
Karl_Felix_Wolff
Major climatic change and biotic turnover during the Triassic
disparity does increase through the Carnian, it increases faster in the Ladinian and Norian, suggesting that the CPE was not a major influence on the rise
Carnian_pluvial_episode
Extinct suborder of reptiles
Scheyer, Torsten M. (2022-07-13). "A new pachypleurosaur from the Early Ladinian Prosanto Formation in the Eastern Alps of Switzerland". Swiss Journal of
Pachypleurosauria
nov Valid Romer Ladinian Chañares Formation Argentina A member of the lagerpetonidae. Lagosuchus fam et gen et sp nov valid Romer Ladinian Chañares Formation
1971_in_paleontology
Extinct family of ray-finned fishes
ray-finned fish, that lived from the Middle Triassic to the Late Jurassic (Ladinian to Tithonian). They are noted for their relatively deep bodies and are
Dapediidae
Extinct genus of reptiles
2003, are junior synonyms of G. tangae. The genus is also known from the Ladinian-aged Middle Triassic Zhuganpo Formation; additionally, the species "Callawayia"
Guizhouichthyosaurus
Extinct family of reptiles
Candelaria from the Santa Maria Formation of Brazil, which lived during the Ladinian stage of the Middle Triassic. The second youngest known owenettid, Ruhuhuaria
Owenettidae
Extinct genus of jawless fishes
Budurovignathus is an extinct genus of conodont. The Ladinian stage of the Middle Triassic is defined by the first appearance of Budurovignathus praehungaricus
Budurovignathus
Extinct genus of molluscs
cephalopods in the order Ceratitida from the middle and upper Triassic (Ladinian and Carnian) of the Alps, named by Mojsisovics (1897). Nannites is the
Nannites
Animals that have evolved aerial locomotion
into broad wings very similar to those of their modern counterparts. The Ladinian genus Potanichthys is the oldest member of this clade, suggesting that
Flying_and_gliding_animals
Extinct genus of reptiles
(2016-11-01). "A new species of Xinpusaurus (Reptilia, Thalattosauria) from the Ladinian (Middle Triassic) of Xingyi, Guizhou, southwestern China". Journal of Vertebrate
Concavispina
Extinct genus of gastropods
Littorinimorpha. These extinct sea snails lived between the Middle Triassic, Ladinian age (about 242 - 237 million years ago) and the Early Cretaceous, Turonian
Purpuroidea
First age of the Early Triassic epoch
; 2005: The Global boundary Stratotype Section and Point (GSSP) of the Ladinian Stage (Middle Triassic) at Bagolino (Southern Alps, Northern Italy) and
Induan
Extinct genus of reptiles
lariosaurine nothosaurid sauropterygian known from the Middle Triassic (Anisian-Ladinian boundary) of Monte San Giorgio, in southern Switzerland and northern Italy
Ceresiosaurus
Extinct genus of reptiles
European Russia from the Bukobay Gorizont. The locality dates back to the Ladinian stage of the Middle Triassic. Additional material has been described from
Jushatyria
Rock formation in the Alps of Europe
the dolomitic mass of Ladinian - Norian age is divided by the Carnian sandstones and shales of Lunz Formation to the Ladinian - Carnian Wetterstein Dolomite
Main_Dolomite
Extinct genus of Triassic fish
Graulia is an extinct genus of lobe-finned fish from the Middle Triassic (Ladinian) of eastern France. It contains a single species, G. branchiodonta, and
Graulia
Extinct genus of amphibians
Trematolestes is an extinct genus of temnospondyl amphibian from the Lower Keuper (Ladinian, Middle Triassic) of southern Germany. It was first named by Rainer R.
Trematolestes
Suborder of insects
followed by Locustavidae and Dzhajloutshellidae from the mid-Triassic (Ladinian age), roughly 242 to 237 million years ago. The group diversified during
Caelifera
Extinct genus of reptiles
extinct genus of nothosaurid from the Middle Triassic (late Anisian to late Ladinian stage) of central and western Europe and China. With a complete specimen
Lariosaurus
Supercontinent from the late Paleozoic to early Mesozoic eras
north-central Atlantic. The first breakup of Pangaea is proposed for the late Ladinian (230 Ma) with the formation of large half grabens. By 201 Ma, the Central
Pangaea
Extinct genus of reptiles
species of Anshunsaurus, A. huangguoshuensis, was named in 1999 from the Ladinian or Carnian age Falang Formation in Guanling County. Although several complete
Anshunsaurus
Extinct genus of reptiles
armored doswelliid archosauriform reptile known from the Middle Triassic (Ladinian stage) of the Erfurt Formation in Germany. The type species, Jaxtasuchus
Jaxtasuchus
Extinct genus of sponges
Guadalupia is an extinct genus of sea sponges. It includes a number of extinct species including: Guadalupia auricula, G. cupulosa, G. ramescens, G. microcamera
Guadalupia
1017/jpa.2025.10105. Vörös, A.; Budai, T.; Bercsényi, M.; Pintér, Z. (2025). "Ladinian ammonoids from the Balaton Highland (Hungary). II. Veszprémfajsz". Földtani
2025_in_paleomalacology
Extinct genus of reptiles
Eschenau quarry is one of the most fossiliferous sites preserving the Ladinian-age Lower Keuper sequence, also known as the Erfurt Formation. The specimens
Polymorphodon
System that relates geologic strata to time
pseudosuchians to survive. ~205.7 Norian ~227.3 Carnian ~237 * Middle Ladinian ~241.464 * Anisian 247 Lower/Early Olenekian 250.8 Induan 251.902 ± 0.024
Geologic_time_scale
Extinct genus of tetrapodomorphs
extinct genus of bystrowianid chroniosuchians from upper Middle Triassic (Ladinian age) deposits of Kupferzell and Vellberg, northern Baden-Württemberg, Germany
Bystrowiella
Rhaeto-Romance language of northeast Italy
distinguish between ladins dolomitiches (Dolomitic Ladinians) and ladins nonejes (Non Valley Ladinians) at the next census. There is no linguistic census
Ladin_language
Genus of jumping bristletails
of a single species, Gigamachilis triassicus, from the Middle Triassic (Ladinian) aged Meride Limestone of Monte San Giorgio, northern Italy. The species
Gigamachilis
Extinct genus of reptiles
known from a fragmentary partial skeleton. This fossil hails from the late Ladinian or early Carnian-age "basal Isalo II beds". As the first lagerpetid found
Kongonaphon
Genus of Triassic ichthyosaur
informally termed the "Upper Saurian Niveau" of Svalbard, spanning from the Ladinian stage of the Middle Triassic to the Carnian stage of the Late Triassic
Besanosaurus
Genus of procolophonoid reptile
extinct genus of procolophonoid stem-reptile known from the Middle Triassic (Ladinian age) Santa Maria Formation of Brazil. The genus contains a single species
Sauropia
Stratigraphic Group in Western Canada
Schooler Creek Group is a stratigraphic unit of Middle to Late Triassic (Ladinian to Norian) age in the Western Canadian Sedimentary Basin. It is present
Schooler_Creek_Group
Extinct genus of reptiles
France. These were found in the upper Muschelkalk, which dates back to the Ladinian stage of the Middle Triassic. Material found in France includes the holotype
Simosaurus
Description Images †Lonchidion †L. selachos †L. griffisi †L. babulskii Ladinian–Maastrichtian †L. selachos: Saskatchewan, Canada; USA †L. griffisi: Wyoming
List of vertebrate fauna of the Campanian stage
List_of_vertebrate_fauna_of_the_Campanian_stage
Extinct genus of aquatic reptiles
cladogram below: Shang, Qing-Hua; Wu, Xiao-Chun; Li, Chun (2020-05-03). "A New Ladinian Nothosauroid (Sauropterygia) from Fuyuan, Yunnan Province, China". Journal
Brevicaudosaurus
Clade of therapsids
Hans-Dieter (June 2006). "A traversodont cynodont from the Middle Triassic (Ladinian) of Baden-Württemberg (Germany)". Paläontologische Zeitschrift. 80 (2):
Cynodontia
Genus of extinct reptiles
(2022). "An unusual Colognathus-like reptile from the Middle Triassic (Ladinian) Erfurt Formation of Germany". Neues Jahrbuch für Geologie und Paläontologie
Quasicolognathus
Extinct genus of cartilaginous fishes
Strophodus is an extinct genus of durophagous hybodont known from the Triassic to Cretaceous. It was formerly confused with Asteracanthus. It has long
Strophodus
Extinct order of reptiles
choristodere remains have also been reported from the Middle Triassic (Ladinian) Erfurt Formation of Germany. However, these remains were suggested to
Choristodera
Extinct family of reptiles
Azendohsaurus itself has been dated from the end of the Middle Triassic during the Ladinian (at least for A. madagaskarensis) into the Carnian stage of the earliest
Azendohsauridae
Extinct genus of reptiles
phylogenetic analysis by Stocker et al. (2017) Diandongosuchus was found in a Ladinian-age marine limestone formation that has preserved many marine reptiles
Diandongosuchus
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