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First stage of the Devonian
The Lochkovian is one of three faunal stages in the Early Devonian Epoch. It lasted from 419.62 ± 1.36 million years ago to 413.02 ± 1.91 million years
Lochkovian
First epoch/series of the Devonian
series. It lasted from 419.62 ± 1.36 to 393.47 ± 0.99 and began with the Lochkovian Stage 419.62 ± 1.36 to 413.02 ± 0.99, which was followed by the Pragian
Early_Devonian
Extinct genus of cartilaginous fishes
fish. It lived from the Pridoli to the Lochkovian. The type species I. gracilis is only known from Lochkovian. Ischnacanthus was first discovered in 1861
Ischnacanthus
Extinct genus of jawless fishes
Ludlow), or Early Devonian (Lochkovian). It was possibly a nectonic filter, of active locomotion, due to its physiognomy. Lochkovian of United Kingdom (Scotland)
Cowielepis
Fourth period of the Paleozoic Era
The Early Devonian lasted from 419.62 to 393.47 Ma. It began with the Lochkovian Stage from 419.62 to 413.02 Ma, which was followed by the Pragian from
Devonian
Genus of millipedes
Scotland during the latest Silurian and earliest Devonian period (Pridoli–Lochkovian epoch). It is one of the first myriapods, and among the oldest creatures
Pneumodesmus
Second stage of the Devonian
years ago to 410.62 ± 1.95 million years ago. It was preceded by the Lochkovian Stage and followed by the Emsian Stage. The most important Lagerstätte
Pragian
Extinct genus of plants
nematophyte with internally thickened tubes, dating from the Wenlock to the Lochkovian epochs. It is thought to be terrestrial or freshwater, and seems to be
Nematasketum
Genus of galeaspid vertebrates
Lungmenshanaspis is an extinct genus of galeaspid vertebrates. It lived during the Lochkovian stage in Sichuan, China. Lungmenshanaspis was 15 cm (5.91 in) in length
Lungmenshanaspis
Extinct genus of lungfish
of prehistoric lungfish which lived during the Early Devonian period (Lochkovian to Pragian stages, about 407-416 million years ago). Fossils of Y. praecursor
Youngolepis
Final Series (Epoch) of the Silurian
Epoch succeeds the Ludfordian age of the Ludlow epoch and precedes the Lochkovian, the earliest of three ages within the Early Devonian geological epoch
Pridoli_Epoch
Genus of extinct jawless fish
species of the genus, C. lyelli was described from the Early Devonian (Lochkovian)-aged Old Red Sandstone of Scotland in 1835. However this species is poorly
Cephalaspis
Extinct genus of Acanthodian fish
Early Ludlow from the Mackenzie Mountains in Canada, X. minutus from Lochkovian of Spitsbergen in Norway, and X. grandis from Siegenian of Spitsbergen
Xylacanthus_(fish)
Extinct genus of scorpion
revision of Praearcturus gigas: a giant scorpion from the Lower Devonian (Lochkovian) of Britain". Palaeontology. 69 (3) e70064. doi:10.1111/pala.70064. ISSN 0031-0239
Praearcturus
Mitraspis is an extinct genus of pteraspidid that lived during the Lochkovian stage of the Early Devonian epoch. Mitraspis cracens is known from the Drake
Mitraspis
Extinct genus of jawless fishes
genus of pteraspidid heterostracan jawless fish. It existed from the Lochkovian to Eifelian epochs of the Devonian period in areas that are now the Americas
Pteraspis
Cambrian era geologic formation in the Southeastern United States
Pragian Oriskany Sandstone Ridgeley Sandstone Wildcat Valley Sandstone Lochkovian Helderberg Group Keyser Limestone Oriskany Sandstone Ridgeley Sandstone
Copper_Ridge_Dolomite
Class of arthropods
likely originated during the Pridoli Epoch of the late Silurian or the Lochkovian of the early Devonian. During the Carboniferous, Arthropleura became the
Millipede
Third period of the Paleozoic Era
Dianne; Wellman, Charles (2001). "Embryophytes on Land: The Ordovician to Lochkovian (Lower Devonian) Record". Plants Invade the Land. pp. 3–28. doi:10.7312/gens11160-003
Silurian
(2002). "Paucicanthus vanelsti gen. et sp. nov., an Early Devonian (Lochkovian) acanthodian that lacks paired fin-spines". Canadian Journal of Earth
List_of_acanthodian_genera
Genus of placoderm fishes
phyllolepid placoderm which lived during the Early Devonian period (Late Lochkovian epoch), of Qujing, Yunnan province, south China. The known fossil remains
Gavinaspis
Geological formation in Yunnan, China
remains of fossilized fish and plants of the Early Devonian period (Late Lochkovian). It was originally referred to as the Xitun Member of the Cuifengshan
Xitun_Formation
Extinct genus of arthropods
to Emsian epoch of Guangxi; and H. yueya, the type species, from the Lochkovian epoch of Yunnan. The name of the genus is derived from the Chinese character
Houia
Extinct genus of ophiurinid echinoderm
Austrophiaulax Temporal range: Lochkovian PreꞒ Ꞓ O S D C P T J K Pg N Scientific classification Kingdom: Animalia Phylum: Echinodermata Family: †Ophiurinidae
Austrophiaulax
Extinct genus of scorpions
Proscorpius Temporal range: Wenlock-Lochkovian, 422.9–416.0 Ma PreꞒ Ꞓ O S D C P T J K Pg N Fossil specimen YPM IP 545850 Life restoration Scientific classification
Proscorpius
System that relates geologic strata to time
47 ± 0.99 * Lower/Early Emsian 410.62 ± 1.95 * Pragian 413.02 ± 1.91 * Lochkovian 419.62 ± 1.36 * Silurian Pridoli Laurentia and Avalonia-Baltica collide
Geologic_time_scale
Devonian sedimentary rocks in South Africa
Bokkeveld Group rocks are considered to range between Lower Devonian (Lochkovian) to Middle Devonian (Givetian) in age. The Cape Supergroup rocks were
Bokkeveld_Group
Geological formation in Ohio
Lower Emsian Amherstburg Dolomite Detroit River Group Hillsboro Sandstone Lochkovian Holland Quarry Shale Bass Islands Dolomite Bois Blanc Formation Helderberg
Kerbel_Formation
Extinct genus of bony fishes
fish which lived from around 418 to 397 million years ago (Pridoli to Lochkovian stages). Fossils of Psarolepis have been found mainly in South China and
Psarolepis
Extinct genus of arthropods
family Stylonuridae. It contains three species, all from the Devonian (Lochkovian to Pragian); P. plotnicki from Nunavut, Canada and P. sturrocki and P
Pagea
Extinct genus of fishes
Diploholcaspis Temporal range: Lower Devonian, Lochkovian PreꞒ Ꞓ O S D C P T J K Pg N Scientific classification Kingdom: Animalia Phylum: Chordata Infraphylum:
Diploholcaspis
Class of chordates
Polybranchiaspidida appear in the fossil record a little after the beginning of the Lochkovian Epoch, at the start of the Devonian. The vast majority of the supraorder's
Galeaspida
Devonian fossil locality in Canada
Delorme Group. An earliest Devonian (Lochkovian) age is established by its fossil fauna, with distinctive Lochkovian fish such as Waengsjoeaspis, Canadapteraspis
MOTH_locality
Rhuddanian Aeronian Telychian Sheinwoodian Homerian Gorstian Ludfordian Lochkovian Pragian Emsian Dent Group Stockdale Group Tranearth Group Coniston
Windermere_Supergroup
Extinct paraphyletic class of fishes
during the Late Silurian, reaching their apex of diversity during the Lochkovian stage of the Early Devonian, declining during the Pragian but rising again
Acanthodii
(2026). "Bataspis crux, a galeaspid (jawless stem gnathostome) from the Lochkovian of Yunnan with a batwing-shaped head adapted for hydrodynamic efficiency"
2026_in_paleoichthyology
Extinct genus of cartilaginous fishes
two species of this genus, both of which lived in the lower Devonian (Lochkovian) of the Northwest Territories, Canada. Hanke, Gavin F.; Davis, Samuel
Tetanopsyrus
First stage of the Silurian
Rhuddanian Aeronian Telychian Sheinwoodian Homerian Gorstian Ludfordian Lochkovian O L Hirnantian ← Lau event ← Mulde event ← Ireviken event Subdivision
Rhuddanian
Genus of Chondrichthyes
was described in the Lochkovian of Ukraine. It is also reported from the Lochkovian and Pragian of Spain, as well as the Lochkovian of the MOTH site and
Altholepis
Widespread geologic group in the Southeastern United States
Lower Emsian Amherstburg Dolomite Detroit River Group Hillsboro Sandstone Lochkovian Holland Quarry Shale Bass Islands Dolomite Bois Blanc Formation Helderberg
Knox_Supergroup
Final stage of the Devonian
– −370 — – −365 — – −360 — – Paleozoic Devonian C M Early Mid Late E Lochkovian Pragian Emsian Eifelian Givetian Frasnian Famennian Tournaisian S Přídolí
Famennian
Extinct genus of jawless vertebrates
Nochelaspis is an extinct genus of galeaspid vertebrates. It lived during the Lochkovian period in Lower Devonian East Asia (in today southeastern China and northern
Nochelaspis
Widespread shallow marine limestone
Lower Emsian Amherstburg Dolomite Detroit River Group Hillsboro Sandstone Lochkovian Holland Quarry Shale Bass Islands Dolomite Bois Blanc Formation Helderberg
Trenton_Group
Embryophyte plants with branched spore-bearing generation
Edwards, D. & Wellman, C. (2001), "Embryophytes on Land: The Ordovician to Lochkovian (Lower Devonian) Record" in Gensel & Edwards 2001, pp. 3–28 Taylor, T
Polysporangiophyte
Extinct genus of jawless fishes
Liliaspis Temporal range: Lochkovian Scientific classification Kingdom: Animalia Phylum: Chordata Infraphylum: Agnatha Class: †Pteraspidomorpha Subclass:
Liliaspis
Large Geologic Group
Lower Emsian Amherstburg Dolomite Detroit River Group Hillsboro Sandstone Lochkovian Holland Quarry Shale Bass Islands Dolomite Bois Blanc Formation Helderberg
Salina_Group
Stratigraphical unit of Upper Ordovician age in the Appalachian Basin
Lower Emsian Needmore Shale Huntersville Chert Pragian Oriskany Sandstone Lochkovian Helderberg Group S Pridoli Bertie Formation Bass Islands Formation Keyser
Utica_Shale
Municipal District of Prague in Czech Republic
southwestern part of the city. It is most notable for being the namesake of the Lochkovian stage. "Archived copy" (PDF). Archived from the original (PDF) on 2024-07-12
Lochkov
Geologic formation in Yunnan, China
and galeaspid fish and it represents the Early Devonian period (Early Lochkovian) of China. A new species of Galeaspida was discovered in the formation
Xishancun_Formation
Extinct genus of fishes
including a marvelously preserved, intact specimen, are known from the Lochkovian Epoch-aged Xitun Formation of Early Devonian Yunnan. The armor is very
Parayunnanolepis
[[Silurian]] aged formation
Formation Helderberg Group (geology) Pragian Helderberg Group (geology) Lochkovian Helderberg Group (geology) Keyser Formation Manlius Limestone S Pridoli
Irondequoit_Formation
Silurian geologic formation in Ghat, Libya
Fossilworks.org Plotnick, 1999 Plotnick, R. E (1999), Habitat of Llandoverian-Lochkovian eurypterids, in A. J. Boucot, J. D. Lawson (eds.), Paleocommunities -
Acacus_Sandstone
Extinct genus of chelicerate
euchelicerate linking horseshoe crabs and eurypterids, from the Lower Devonian (Lochkovian) of Yunnan, China". Zoologica Scripta. 44 (6): 645–652. doi:10.1111/zsc
Bembicosoma
those of the acanthothoracid, Connemarraspis, have been found in the late Lochkovian-aged marine strata of the Connemarra Formation in Australia. It is considered
Narrominaspis
Genus of extinct fish
is an extinct monospecific genus of arthrodire placoderm fish found in Lochkovian-Pragian (Lower Devonian) deposits of Podolia, Western Ukraine. The type
Erikaspis
Fossilised primitive plant spore
abrupt decline in diversity and abundance around 410 Ma during the latest Lochkovian (Early Devonian), with only a few forms persisting into the Emsian period
Cryptospore
Class of lobe-finned fishes
Diabolepis, dating to 419–417 million years ago, during Early Devonian (Lochkovian), are the currently oldest known lungfish, and show that the lungfishes
Lungfish
Geological formation in Ohio
Lower Emsian Amherstburg Dolomite Detroit River Group Hillsboro Sandstone Lochkovian Holland Quarry Shale Bass Islands Dolomite Bois Blanc Formation Helderberg
Cedarville_Dolomite
Paraphyletic infraphylum of fish
and distribution of associated conodonts from the Llandovery-lowermost Lochkovian of the Welsh Borderland". Palaeontology. 47 (5): 1211–1265. Bibcode:2004Palgy
Agnatha
Extinct genus of jawless fishes
Tesseraspis Temporal range: Lochkovian PreꞒ Ꞓ O S D C P T J K Pg N Scientific classification Kingdom: Animalia Phylum: Chordata Infraphylum: Agnatha Class:
Tesseraspis
Extinct genus of fishes
contains a single species, D. speratus of Yunnan, China, from the mid-late Lochkovian of the Xitun Formation. It is one of the oldest known lungfish genera
Diabolepis
Extinct genus of millipedes
formation it contains actually appears to be from the Early Devonian (Lochkovian) instead. Although a 2023 study confirmed the age identification of the
Cowiedesmus
Extinct genus of euaxosid echinoderm
Acruxaster Temporal range: Ludlow–Lochkovian PreꞒ Ꞓ O S D C P T J K Pg N Scientific classification Kingdom: Animalia Phylum: Echinodermata Class: Asteroidea
Acruxaster
Geological formation in Scotland
Wenlock to Ludlow, but zircon geochronology points to the Early Devonian, Lochkovian instead. In study published in 2023, according to spore microfossils and
Cowie_Formation
Chronological outline of major events in the development of plants
D.; Wellman, C.H. (2001). "2. Embryophytes on Land: The Ordovician to Lochkovian (Lower Devonian) Record". Plants Invade the Land. Columbia University
Timeline_of_plant_evolution
Order of arthropods
euchelicerate linking horseshoe crabs and eurypterids, from the Lower Devonian (Lochkovian) of Yunnan, China". Zoologica Scripta. 44 (6): 645–652. doi:10.1111/zsc
Chasmataspidida
Lithostratigraphic unit
Shale Needmore Shale Oriskany Sandstone Shriver Chert Pragian Corriganville Limestone Helderberg Formation Lochkovian Keyser Limestone New Creek Limestone
Greenbrier_Group
Extinct genus of fishes
sarcopterygian, lobe-finned fish which lived during the Early Devonian (Lochkovian) period of East Yunnan, South China. Because it is clear that the jaws
Styloichthys
Geologic Group found in the Appalachian Basin
Helderberg Group Stratigraphic range: Lochkovian-Emsian ~419–393 Ma PreꞒ Ꞓ O S D C P T J K Pg N Type Group Sub-units Port Ewen Shale Minisink Limestone
Helderberg_Group_(geology)
Second epoch of the Devonian period
−370 — – −365 — – −360 — – Paleozoic Devonian C M Early Mid Late Early M Lochkovian Pragian Emsian Eifelian Givetian Frasnian Famennian S Přídolí ← Hangenberg
Middle_Devonian
The Becraft Formation is a geologic formation of marine sedimentary rock found in New York State. The Becraft is a part of the lower Devonian Helderberg
Becraft_Formation
Family of brachiopods
dropped, but they slowly recovered until their demise at the end of the Lochkovian of the Early Devonian. Orthids are known from many regions, including
Orthidae
Geologic Unit found in the Appalachian Basin
Shale Needmore Shale Oriskany Sandstone Shriver Chert Pragian Corriganville Limestone Helderberg Formation Lochkovian Keyser Limestone New Creek Limestone
Tully_Formation
Extinct genus of cephalopods
Lorieroceras Temporal range: Lochkovian-Pragian Reconstruction of L. lorieri Scientific classification Kingdom: Animalia Phylum: Mollusca Class: Cephalopoda
Lorieroceras
Extinct genus of fishes
Zhanjilepis aspartilis is an extinct yunnanolepid placoderm from Pridoli and Lochkovian rocks of Hunan, China. Zhanjilepis is known only from distinctively ordered
Zhanjilepis
Extinct genus of crustaceans
Warneticaris Temporal range: Telychian-Lochkovian, 439-411 Ma PreꞒ Ꞓ O S D C P T J K Pg N Scientific classification Kingdom: Animalia Phylum: Arthropoda
Warneticaris
Extinct genus of trilobites
Viaphacops Temporal range: Lochkovian-Givetian ~388–383 Ma PreꞒ Ꞓ O S D C P T J K Pg N Viaphacops multicinctus, derived intact length 45mm,↑lateral, ↓dorsal
Viaphacops
Early Devonian genus of placoderm fish
placoderms which lived in shallow marine environments in the early Devonian (Lochkovian). The name Romundina honors Canadian geologist and paleontologist Dr.
Romundina
Extinct genus of jawless fish
The C.grossi species is thought to be from the Devonian Period (Early Lochkovian). Fossils of C.grossi have been found in the Boothia Peninsula. Isolated
Canonia
Fifth stage of the devonian and second stage of the Middle Devonian
– −370 — – −365 — – −360 — – Paleozoic Devonian C M Early Mid Late E Lochkovian Pragian Emsian Eifelian Givetian Frasnian Famennian Tournaisian S Přídolí
Givetian
Extinct genus of chelicerate
euchelicerate linking horseshoe crabs and eurypterids, from the Lower Devonian (Lochkovian) of Yunnan, China". Zoologica Scripta. 44 (6): 645–652. doi:10.1111/zsc
Legrandella
2023. Slavík, Ladislav; Hladil, Jindřich (1 July 2020). "Early Devonian (Lochkovian – early Emsian) bioevents and conodont response in the Prague Synform
Šilalė_Event
Extinct genus of jawless fishes
Drepanaspis (from Greek: δρεπάνη drepánē 'sickle' and Greek: ἀσπίς aspís 'shield') is an extinct genus of heterostracan armoured jawless fish from the
Drepanaspis
Extinct genus of brittle stars
Furada Formation covered partly by vegetation, which dates to the early Lochkovian stage of the Devonian period. In 2015, paleontologists Daniel Blake, Samuel
Ophiocantabria
Extinct genus of cartilaginous fishes
Ptomacanthus Temporal range: Early Devonian (Lochkovian) PreꞒ Ꞓ O S D C P T J K Pg N Scientific classification Kingdom: Animalia Phylum: Chordata Class:
Ptomacanthus
ipuensis. Cladochonus isaacmariai Sp. nov Valid Ernst & May Devonian (Lochkovian) Birdsong Shale United States ( Tennessee) A tabulate coral belonging
2025_in_paleontology
Extinct genus of eurypterid
in length. The first fossils of Pterygotus were found in deposits of Lochkovian-Pragian (Early Devonian) age by quarrymen in Scotland and western England
Pterygotus
Devonian geologic unit in the Appalachian basin
Formation Helderberg Group (geology) Pragian Helderberg Group (geology) Lochkovian Helderberg Group (geology) Keyser Formation Manlius Limestone S Pridoli
Genesee_Formation
Extinct genus of cartilaginous fishes
extinct genus of spiny shark. This genus is known from the Early Devonian (Lochkovian) of Europe, previously considered Silurian remains actually belong to
Climatius
Geological formation in Great Britain
Dartmouth Group Stratigraphic range: Lochkovian to Pragian (Devonian) Rocks of the Dartmouth group at Sugary Cove near Dartmouth Type Group Sub-units Whitsand
Dartmouth_Group
Lower Emsian Needmore Shale Huntersville Chert Pragian Oriskany Sandstone Lochkovian Helderberg Group S Pridoli Bertie Formation Bass Islands Formation Keyser
Greenland_Gap_Group
First age of the Late Devonian epoch
– −370 — – −365 — – −360 — – Paleozoic Devonian C M Early Mid Late E Lochkovian Pragian Emsian Eifelian Givetian Frasnian Famennian Tournaisian S Přídolí
Frasnian
Order of brachiopods
known representatives of the order. It is known from the Lower Devonian (Lochkovian) to the Carboniferous from many places in the world. More than a hundred
Spiriferinida
Geologic formation in Bolivia and Argentina
Paleontology, 47 (3): 549–550 Plotnick, R. E (1999), Habitat of Llandoverian-Lochkovian eurypterids, in A. J. Boucot, J. D. Lawson (eds.), Paleocommunities -
Lipeón_Formation
Extinct genus of chelicerate
euchelicerate linking horseshoe crabs and eurypterids, from the Lower Devonian (Lochkovian) of Yunnan, China". Zoologica Scripta. 44 (6): 645–652. doi:10.1111/zsc
Cyamocephalus
Geologic formation in the United States
Pragian Oriskany Sandstone Ridgeley Sandstone Wildcat Valley Sandstone Lochkovian Helderberg Group Keyser Limestone Oriskany Sandstone Ridgeley Sandstone
Antietam_Formation
Mapped bedrock unit
Shale Needmore Shale Oriskany Sandstone Shriver Chert Pragian Corriganville Limestone Helderberg Formation Lochkovian Keyser Limestone New Creek Limestone
Mauch_Chunk_Formation
{{{type-species}}} {{{type-authority}}}, {{{type-authority-year}}} Latest Silurian(?)–Lochkovian United States Archipelepis Märss, 2001 Type species: A. turbinata Soehn
List_of_thelodont_genera
Extinct genus of trilobites
Trimerus Temporal range: Llandovery–Lochkovian PreꞒ Ꞓ O S D C P T J K Pg N Fossil of Trimerus (Trimerus) delphinocephalus from the Rochester Shale of North
Trimerus
Extinct genus of fishes
averaging up to about 2 centimetres (0.79 in) long, which lived during the Lochkovian epoch in Early Devonian Yunnan Province, China and northern Vietnam. Anatomically
Minicrania
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