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Proteinaceous hair-like appendage on the surface of bacteria
A pilus (Latin for 'hair'; pl.: pili) is a hair-like cell-surface appendage found on many bacteria and archaea. The terms pilus and fimbria (Latin for
Pilus
Topics referred to by the same term
Momtaz, Bangladeshi singer Pilu, Arad, Romania Pilu, Estonia Pilu river, Myanmar Pilu (raga), Hindustani music Pilu oil Pīlu Pilu (TV series), an Indian television
Pilu
Roman military rank
Empire, the emperor Claudius created the office of primus pilus iterum. To become the primus pilus iterum an officer must have formerly served as a tribune
Primus_pilus
Largest military unit of the Roman army
legion was in battle formation, the pilus prior was given command of their entire cohort. The primus pilus was also a pilus prior, and the most senior of all
Roman_legion
Musical artist
"Izwan Pilus meninggal dunia" [Izwan Pilus died]. Utusan (in Malay). 21 January 2012. Izwan Pilus meninggal dunia, Sinar Harian, Singer Izwan Pilus buried
Izwan_Pilus
Punjabi poet (1580–1675)
Pilu (c. 1580–1675) was a Punjabi dramatic poet who is known for his rendition of the Punjabi tragic romance Mirza Sahiban. He was a contemporary of Mughal
Pilu_(poet)
Index of plants with the same common name
Pīlu (from Sanskrit पीलु pīlu) is the common name of two species of tree (Careya arborea orﾠ Salvadora persica). Mentioned in Mahābhārata. The name may
Pīlu
Army officer in Imperial Rome
Castrorum and the Tribunus Laticlavius. The Primus Pilus, being the Pilus Prior of his cohort, did not have a Pilus Posterior, making the first cohort 5 double
Centurion
Genus of gastropods
Pilus is a genus of sea snails, marine gastropod mollusks in the family Bathysciadiidae. Species within the genus Pilus include: Pilus conicus (Verrill
Pilus_(gastropod)
Musical artist
Shahbaz Khan Pilu (born 15 December 1960), commonly known as Pilu Khan, is a Bangladeshi singer, music composer, drummer and director. He is a current
Pilu_Khan
Roman military subdivision
hastatus prior, princeps posterior, princeps prior, pilus posterior, and pilus prior. The pilus prior would command the cohort overall.[citation needed]
Cohort_(military_unit)
Method of bacterial gene transfer
cell produces the pilus (typically a type IV secretion system); necessary genes are expressed from the conjugative plasmid. Pilus attaches to recipient
Bacterial_conjugation
Raga in Hindustani music
Pilu or Peelu is a raga of Hindustani classical music. It is mostly used in light-classical forms, like thumris. 'P 'N S g m P N S' 'N S G m P N S' S g
Pilu_(raga)
periplasmic and extracellular pilus subunits, dimeric usher outer membrane pore and associated Sec machinery. The Pilus subunits polymerise via a non-covalent
Chaperone-usher_fimbriae
infantryman in the Roman Empire. Pilus Prior – Senior centurion of a cohort. Pilus Posterior – Deputy to the pilus prior. Praefectus Castrorum – Camp
List_of_Roman_army_unit_types
Protein family
Pilin refers to proteins that create pilus structures in bacteria. These structures can be used for the exchange of genetic material, or as a cell adhesion
Pilin
Species of gastropod
Pilus conicus is a species of sea snail, a marine gastropod mollusk in the family Pseudococculinidae. The size of the shell is 1 mm. This species occurs
Pilus_conicus
Oil extracted from Pilu tree seeds
Pilu oil is an extract from seeds of the Pilu tree (Salvadora persica), also known as the toothbrush tree. It is used for soaps, detergents, and resist
Pilu_oil
Malaysian singer, actor, dancer and TV host
fellow artist Izwan Pilus. Aril had hosted some tv program long before he drafted to AF. It was when he little bit known as Ajam Pilus. Later he appeared
Aril_(entertainer)
Centurions of the Roman army
J.A. Turney. Pullo is the senior centurion (Primus Pilus) of the XIII while Vorenus was the Pilus Prior (Centurion of cohort his first century is attached)
Vorenus_and_Pullo
Form of crawling bacterial motility
The type IV pilus complex consists of both the pilus itself and the machinery required for its construction and motor activity. The pilus filament is
Twitching_motility
Species of bacterium
extension and retraction of the type IV pilus, respectively. The adhesive functions of the gonococcal pilus play a role in microcolony aggregation and
Neisseria_gonorrhoeae
Commune in Arad, Romania
Pilu (Hungarian: Nagypél) is a commune in Arad County, Romania, is situated on the Criș Plateau, in the north-western part of the county, at the western
Pilu,_Arad
Type of skullcap worn by clergy
denominations of Lutheranism, Anglicanism, and Methodism. It is also called a pilus, pilos, pileus, pileolo, subbiretum, submitrale, soli deo, berrettino, calotte
Zucchetto
B, Rodrigues C, et al. (January 2016). "Structure of a Chaperone-Usher Pilus Reveals the Molecular Basis of Rod Uncoiling". Cell. 164 (1–2): 269–278
P_fimbriae
Bacterium, genetic engineering tool
IV secretion system (T4SS). The structure of the T-pilus showed that the central channel of the pilus is too narrow to allow the transfer of the folded
Agrobacterium_tumefaciens
2022 Indian television series
Pilu is a 2022 Indian Bengali drama television series that released on 10 January 2022 on Zee Bangla. It is also available on the digital platform ZEE5
Pilu_(TV_series)
Musical artist
Pilu Momtaz (c. 1953 – 23 May 2011) was a Bangladeshi singer. Momtaz was born in Dhaka. She was the third of seven children born to the Bangladeshi singer
Pilu_Momtaz
Protein complex found in bacteria and archaea
well characterized. The pilus is composed of VirB2 and VirB5, with VirB2 being the major component. In A. tumefaciens, the pilus is 8-12 nm in diameter
Type_IV_secretion_system
nanomachines involved in bacterial secretion with particular emphasis on pilus biogenesis by the Chaperone-Usher pathway and on Type IV Secretion (T4S)
Gabriel_Waksman
2nd Century AD Roman military commander
leg(ionis) | VI Victric(is)|[...] Translated: Lucius Artorius Castus, Primus Pilus of the legion V Macedonica, Prefect of the Legion VI Victrix [....] An undated
Lucius_Artorius_Castus
Roman soldier
Caelius (c. March 45 BC – c. September AD 9) was the senior centurion (Primus pilus) in XVIII Roman Legion who was killed in the Battle of the Teutoburg Forest
Marcus_Caelius
Biological process of breaking down food
Schematic drawing of bacterial conjugation. 1- Donor cell produces pilus. 2- Pilus attaches to recipient cell, bringing the two cells together. 3- The
Digestion
Body hair removal
the Latin "depilare," a word formed from the prefix "de-" and the root "pilus," meaning "hair." For centuries, hair removal has long shaped gender roles
Hair_removal
Type of bacterium
factor sequence, and do not become F + due to its inability to form a sex pilus. In conjugation mediated by Hfr cells, transfer of DNA starts at the origin
Hfr_cell
Medical condition
Pili multigemini Other names Pili gemini An extracted pilus multigeminus with three distinct hairs growing from the same source. Specialty Dermatology
Pili_multigemini
Species of pathogenic bacterium found in water
V. vulnificus has genes that encode for the biosynthesis of a type IV pilus. These genes have been located on the large chromosome and may help V. vulnificus
Vibrio_vulnificus
Group of prokaryotic enzymes
S2CID 25384605. Swierczynski A, Ton-That H (September 2006). "Type III pilus of corynebacteria: Pilus length is determined by the level of its major pilin subunit"
Sortase
Cīrava castle. p. 1. Zarāns p. 28. Latvijas Piļu un Muižu asociācija. "Kalnamuiža (Sieksātes) muiža". Latvijas Piļu un Muižu asociācija. Archived from the
List of palaces and manor houses in Latvia
List_of_palaces_and_manor_houses_in_Latvia
Bodyguards of the Roman emperors
centurions in one or several legions before achieving the status of Primus pilus (the highest ranked Centurion in a legion). Upon return to Rome, they occupied
Praetorian_Guard
Specialized subunit within a cell
protrudes outside the cell, and its motor, as well as the largely extracellular pilus, are often spoken of as organelles. In biology, organs are defined as confined
Organelle
for DNA in bacterial conjugation. A mating bridge is different from a sex pilus, which is a structure made by an F+ strain bacterium in bacterial conjugation
Mating_bridge
1958 Indian film
Bhosle Pilu (raga) "Barso Re Bairi Badarwa" Asha Bhosle Pilu (raga) "Shokh Shokh Ankhen" Asha Bhosle Pilu (raga) "Sun Ja Pukar" Asha Bhosle Pilu (raga)
Phagun_(1958_film)
Ability of bacteria to move independently using metabolic energy
counterclockwise (CCW). A pilus (Latin for 'hair') is a hair-like appendage found on the surface of many bacteria and archaea. The terms pilus and fimbria (Latin
Bacterial_motility
Musical artist
(Tukaram) – in Raga Jhinjhoti Sharana Sharana – Abhang (Sant Tukaram) – in Raga Pilu. Dhanya Dhanya – Abhang (Sant Eknath) – in Raga Durga. Sada Majzhe – Abhang
Ranjani–Gayatri
Roman Empire honorable discharge from military service
sesterces for the common soldier and around 600,000 sesterces for the primus pilus until the Principate of Caracalla. The honorably discharged legionary also
Honesta_missio
Janya raga of Carnatic music
the Hindustani raag and thaat Kafi. The equivalent raag in Hindustani is Pilu. Kāpi is an audava-vakra sampoorna rāgam with an ascending pentatonic scale
Kapi_(raga)
Element of hierarchy in armed forces
centurions was called primus pilus. The ranks of centurions in the individual cohorts were, in descending order, pilus prior, pilus posterior, princeps prior
Military_rank
Transfer of genes from unrelated organisms
that will be moved to the other bacterium 4: Pilus and T4SS. Conjugation in bacteria using a sex pilus; then the bacterium that received the plasmid
Horizontal_gene_transfer
It is known chiefly from a single individual, Titus Balventius, a primus pilus in the command of Quintus Titurius Sabinus in Gaul. He was severely wounded
Balventia_gens
Upright loops, tufts, or strands of yarn extending from the ground of a fabric
plush, and Turkish towels (terrycloth). The word is derived from Latin pilus for "hair". The surface and the yarn in these fabrics are also called "pile"
Pile_(textile)
Roman Empire from about 27 BC to 476 AD
(centurio primus pilus). Senior centurions included those in command of the five centuriae in the 1st Cohort and the centuriones pilus prior ("front-spear")
Imperial_Roman_army
Group of Roman emperors
of humble origin who had risen through the ranks to the post of primus pilus (which also entailed admission to the equestrian order) were placed as heads
Illyrian_emperors
Haungtharaw River Yunzalin River Moei River (Thaungyin River) Pawn River Pilu River Pai River Teng River Hsim River Nam Pang River Nanding River Ye River
List_of_rivers_of_Myanmar
2025 film directed by Shen Ao
Wang Jingwei's collaborationist government's guard brigade stationed at Pilu Temple [zh], where he hid the album in a restroom. In 1941, the album was
Dead_to_Rights_(film)
Domain of microorganisms
structure called the type IV pilus, and gliding motility, that uses other mechanisms. In twitching motility, the rod-like pilus extends out from the cell
Bacteria
High-ranking Roman military officer
Tribunus laticlavius, Praefectus castrorum, Tribunus angusticlavii & Primus pilus who would collectively be responsible for the legion's operational effectiveness
Legate_(ancient_Rome)
1984 film by Prakash Mehra
Pyar De" (Male) Kishore Kumar Pilu (raga) "Mujhe Naulakha Manga De Re, O Saiyaan Deewane" Kishore Kumar, Asha Bhosle Pilu (raga) "Inteha Ho Gayi Intezar
Sharaabi
Circular plasmid used in creation of transgenic plants
CA, Aguilar J, Ward D, Zambryski P (September 2007). "VirB1* promotes T-pilus formation in the vir-Type IV secretion system of Agrobacterium tumefaciens"
Ti_plasmid
Village in Shan State, Myanmar
a roadway. The Mobye Dam, located within the village, is built along the Pilu River (also called the Baluchaung River). Together with the Lawpita Dam,
Mobye
2016 film by Rakeysh Omprakash Mehra
Directed by Rakeysh Omprakash Mehra Written by Gulzar Based on Mirza Sahiban by Pilu Produced by Rohit Khattar Rakeysh Omprakash Mehra P. S. Bharathi Rajiv Tandon
Mirzya_(film)
Indian actress
Romi Meitei 2008 Malla Malla Leinungshi Usha Amar Raj Kangla Karabar Dasini Pilu H. Kekoo Lotpee Lawyer Homen D' Wai Handakta Nouna Khangnaba Nganthoi P.
Kamala_Saikhom
Indian actress and model
Year Title Director Studio 2012 Ang Tamo Pilu Heigrujam Pilu Films 2015 It's Not My Choice Priyakanta Laishram Priyakanta Productions 2021 Apaiba Leichil
Bishesh_Huirem
Roman legion
Valerianus legatus legionis ? 227-? 229 Sextus Julius Possessorus primus pilus last quarter 1st century Cappadocia CIL II, 1180 Gaius Caristanius Fronto
Legio_XII_Fulminata
Indian actor and singer from Manipur
Thanil Ajit Ningthouja 2018 Soinairaba Thamoi Paikhomba Pilu H. Leichilda Pallaba Thaja Malangba Pilu H. 2022 Awaiba Mapu Ibothoi Khoibam Homeshwori
Bonium_Thokchom
2018 Indian Bengali TV series
roles. After 3 years, the show went ended on 9 January 2022 to make way for Pilu. The story revolves around Shyama, a dark-skinned girl, who sets out to remove
Krishnakoli
Roman legion
angusticlavius Aegyptus AD 69-79 CIL 10, 06976 Lucius Decrius Longinus Primus pilus and Praefectus castrorum Aegyptus AD 101-140 AE 1913, 00215 Tiberius Claudius
Legio_XXII_Deiotariana
Village in Estonia
Pilu is a village in Põltsamaa Parish, Jõgeva County in eastern Estonia. "X-GIS(4) Portal". xgis.maaamet.ee. Retrieved 26 July 2021. 58°42′42″N 25°48′30″E
Pilu,_Estonia
Season of television series
was closed. Bottom two: Claudia Fay Geres & Khairil Azam Pilus Eliminated: Khairil Azam Pilus Special judge: Adibah Noor Special guests: Najwa & Fazura
Akademi_Fantasia_season_7
Biological process
between two bacteria mediated by an external appendage called the conjugation pilus. Bacterial conjugation is controlled by plasmid genes that are adapted for
Sexual_reproduction
One of the ranks of Roman centurions
Each of the ten cohorts that made up a legion had at its head the rank of pilus prior followed by the princeps prior. The princepi priori were often titled
Princeps_prior
Malaysian actor and singer
September 2025. Yusoff, Nor (14 September 2024). "Hari Jadi Fattah, Netizen Pilu!" [Fattah's birthday, netizens are sad!]. Pa&Ma (in Malay). Retrieved 22
Fattah_Amin
Mark and tool of the centurion (Ancient Rome)
The epitaph of M. Caelius, chief centurion (primus pilus) of the ill-fated 18th Legion. His vine staff breaks the frame and even runs across the inscription
Vine_staff
School in Damansara Heights, Kuala Lumpur
Bintang in Kuala Lumpur, Malaysia. The founder and mentor, Dato' Freida Pilus, started it to provide the best possible education for her sons. Over the
Cempaka_Schools
Perennial plant with blue berries
Kusum Mafura Mahua Mango butter Murumuru Nagkesar Palm (kernel) Phulwara Pilu Pongamia Sal-seed (Shorea robusta) Sandalwood Shea butter Tamanu Tea-seed
Blueberry
Aromatic resin from the Commiphora myrrha tree
Kusum Mafura Mahua Mango butter Murumuru Nagkesar Palm (kernel) Phulwara Pilu Pongamia Sal-seed (Shorea robusta) Sandalwood Shea butter Tamanu Tea-seed
Myrrh
Manor house in Latvia
has media related to Jaunmokas Manor. Latvijas Piļu un Muižu asociācija. "Jaunmokas Manor". Latvijas Piļu un Muižu asociācija. Archived from the original
Jaunmokas_Manor
2013 Indian film
Ureinung is a 2013 Manipuri film directed by Pilu Heigrujam and produced by Bandana Maisnam and Ilu, under the banner of Bandana films. It stars Sonia
Ureinung
Topics referred to by the same term
suite TCP (antiseptic) Tenocyclidine, an anesthetic drug Toxin-coregulated pilus, a protein that allows Vibrio cholerae to adhere to enterocytes Transcutaneous
TCP
Punjabi Sufi poet (1658–1707)
Punjab, Pakistan. After Pilu, the poet, he was the second one to write the folktale of Mirza Sahiban as he was asked by the Pilu in a dramatic way to write
Hafiz_Barkhurdar
North American carbonated beverage
Kusum Mafura Mahua Mango butter Murumuru Nagkesar Palm (kernel) Phulwara Pilu Pongamia Sal-seed (Shorea robusta) Sandalwood Shea butter Tamanu Tea-seed
Root_beer
Species of flowering plant in the family Amaryllidaceae
Kusum Mafura Mahua Mango butter Murumuru Nagkesar Palm (kernel) Phulwara Pilu Pongamia Sal-seed (Shorea robusta) Sandalwood Shea butter Tamanu Tea-seed
Allium_ursinum
Salix, genus of trees
Kusum Mafura Mahua Mango butter Murumuru Nagkesar Palm (kernel) Phulwara Pilu Pongamia Sal-seed (Shorea robusta) Sandalwood Shea butter Tamanu Tea-seed
Willow
Italian computer scientist
Pierluigi "Pilu" Crescenzi is a full professor of computer science at the Gran Sasso Science Institute. His research areas include theoretical computer
Pierluigi_Crescenzi
Series of historical novels by Simon Scarrow
a legion in Egypt, with Macro receiving a temporary promotion to Primus pilus (senior centurion). Macro has been close friends with Cato since Cato saved
Eagles_of_the_Empire
100-man military unit in Ancient Rome
the first cohort, who was referred to as primus pilus (the name denoted the first maniple, Latin: pilus, of the first cohort). The term centuria was later
Centuria
Indonesian singer and songwriter (born 1987)
"Pilu Membiru", a track which addressed about loss and grief, was picked as the album's final single on 13 November 2019. The music video, titled Pilu
Kunto_Aji
Roman emperor in 238
Marcellina (otherwise unknown), and who started his career as a Centurio primus pilus before becoming a tribunus militum, and then a praetor. Pupienus's career
Pupienus
Musical scale in Hindustani classical music
is the same as the harmonic minor in western music. There are shades of Pilu in Kirwani. This raga is said to have been borrowed from the Carnatic music
Kirwani
Watchmen of ancient Rome
then transfer to the Legions, where they could reach the rank of Primus Pilus. The prefect was also known as the Prefect of the Watch. The prefect was
Vigiles
Foods called nuts, which may be true nuts or seeds, drupes or legumes
Kusum Mafura Mahua Mango butter Murumuru Nagkesar Palm (kernel) Phulwara Pilu Pongamia Sal-seed (Shorea robusta) Sandalwood Shea butter Tamanu Tea-seed
Nut_(food)
Species of mushrooms
Kusum Mafura Mahua Mango butter Murumuru Nagkesar Palm (kernel) Phulwara Pilu Pongamia Sal-seed (Shorea robusta) Sandalwood Shea butter Tamanu Tea-seed
Matsutake
Chong, Zhang Liangcai, Lu Chaohai, Chen Yueduan, Chen Wencai, Chen Jie, Chen Pilu, Chen Ke, Fan Kaiyou, Fan Yuquan, Yan Shuai, Yan Xiangkan, Luo Guiyou, He
Yunnan delegation to the National People's Congress
Yunnan_delegation_to_the_National_People's_Congress
Topics referred to by the same term
also refer to: Fimbria (bivalve), a genus of clams Fimbria, also called Pilus, hair-like projection on bacteria and archaea Fimbria (female reproductive
Fimbria
Indian actress
acted as a supporting character in 2022 Bengali-language Indian TV series Pilu. Paul made her silver screen debut with Himel Ashraf's 2023 film Priyotoma
Idhika_Paul
Roman legion
fulfillment of a vow, by Lucius Latinius Macer, who describes himself as primus pilus (chief centurion) and as praefectus castrorum ("prefect of the camp", i
Legio_IX_Hispana
Sequence of bacterial DNA
in the recipient cell. tra-region (transfer genes): Genes coding the F-Pilus and DNA transfer process. IS (Insertion Elements) composed of one copy of
F-plasmid
Species of tree
Kusum Mafura Mahua Mango butter Murumuru Nagkesar Palm (kernel) Phulwara Pilu Pongamia Sal-seed (Shorea robusta) Sandalwood Shea butter Tamanu Tea-seed
Durio_zibethinus
Carbonated soft drink
Kusum Mafura Mahua Mango butter Murumuru Nagkesar Palm (kernel) Phulwara Pilu Pongamia Sal-seed (Shorea robusta) Sandalwood Shea butter Tamanu Tea-seed
Birch_beer
Protein family
The fimbrial usher protein is involved in biogenesis of the pilus in Gram-negative bacteria. The biogenesis of some fimbriae (or pili) requires a two-component
Fimbrial_usher_protein
PILUS
PILUS
PILUS
PILUS
Girl/Female
British, English
Noble Friend
Male
Hungarian
Hungarian form of German Hrodebert, RÓBERT means "bright fame."Â
Girl/Female
Australian, Polish
Truthful; Noble
Boy/Male
Celtic Irish
Oath.
Boy/Male
Indian, Malayalam, Tamil
God of Dev; Graceful
Girl/Female
Greek
Born of beauty.
Surname or Lastname
English
English : apparently a variant spelling of Russon.
Girl/Female
Indian
Prayer; Good Person
Girl/Female
Indian, Marathi
Well Natured; Dark Skinned
Boy/Male
Hindu, Indian, Marathi
Rays of Light
PILUS
PILUS
PILUS
PILUS
PILUS