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North American faunal stage according to the North American Land Mammal Ages chronology
The Hemphillian North American Stage on the geologic timescale is a North American faunal stage according to the North American Land Mammal Ages chronology
Hemphillian
Extinct genus of mammals
genus of camel (tribe Camelini), endemic to North America from the late Hemphillian stage of the Miocene through the Irvingtonian stage of the Pleistocene
Titanotylopus
Extinct genus of proboscideans
M. matthewi is recorded from the late Hemphillian to early Blancan stages. Mammutid specimens of the Hemphillian and Blancan had typically previously been
Mastodon
Extinct genus of mammals
is an extinct equid which was endemic to North America from the late Hemphillian stage of the Miocene through the Zanclean stage of the Pliocene (10.3—3
Dinohippus
Extinct genus of mammals
Ilingoceros Temporal range: Hemphillian Reconstructed horn cores of I. alexandrae, American Museum of Natural History Scientific classification Kingdom:
Ilingoceros
Extinct genus of mammals
to the end of Hemphillian faunal stage from around 17.5 to 4.9 million years ago, and is considered an index fauna of the Hemphillian. However, several
Teleoceras
Second epoch of the Neogene Period
Pliocene. In the system of North American Land Mammal Ages (NALMA) include Hemphillian (9–4.75 Ma), and Blancan (4.75–1.6 Ma). The Blancan extends forward into
Pliocene
Genus of carnivores
"A New Species of Raccoon, Procyon garberi, from Late Miocene (Late Hemphillian) Mehrten Formation, Stanislaus County, California". Bulletin of the Southern
Procyon_(genus)
Extinct genus of bears
width of M1 with a longer talonid than in P. edensis. Likely part of the Hemphillian 2 migration wave (Hh2) across the Beringinian land bridge in the late
Plionarctos
Extinct species of rodent
?Oryzomys pliocaenicus is a fossil rodent from the Hemphillian (late Miocene) of Kansas, central United States. It is known from a single mandible (lower
?Oryzomys_pliocaenicus
Sixth and last age of the Miocene Epoch, about 7 to 5 million years ago
South American Land Mammal Ages, and falls inside the more extensive Hemphillian North American Land Mammal Age. During the Messinian, around 6 million
Messinian
Extinct genus of mammals
dating from the Barstovian in the Miocene to the youngest dating in the Hemphillian faunal stage of the early Pliocene. Based on study of the tooth morphology
Astrohippus
Extinct genus of proboscid
(5.9 to 2.2 Ma). This temporal range also corresponds from the late Hemphillian faunal stage (Hh4) to the Late Blancan faunal stage, with Rhynchotherium
Rhynchotherium
Extinct species of mammal
Creek, Nebraska, Hemphillian (10.3 - 4.9 Ma), Ochotona cf. spanglei Mailbox Prospect, Antelope County, Nebraska, Late/Upper Hemphillian (10.3 - 4.9 Ma)
Ochotona_spanglei
Earliest age on the geologic time scale of the Pliocene era
Tabianian or Dacian of Central Europe. It also corresponds to the late Hemphillian to mid-Blancan North American Land Mammal Ages. In California, the Zanclean
Zanclean
Geologic formation in Florida, United States
Miocene to Early Pliocene North American land mammal age: Barstovian to Hemphillian The Bone Valley Formation includes a diverse assemblage of vertebrate
Bone_Valley_Formation
Genus of birds
are of unclear affinities. An undescribed fossil species from the late Hemphillian (5.0–4.1 mya) of Jalisco, central Mexico, has also been identified from
Pygmy_goose
Genus of mammal
Tayassu Temporal range: Late Hemphillian to recent, 9.4–0 Ma PreꞒ Ꞓ O S D C P T J K Pg N Possible not described Early Miocene records of cf. Dicotyles[citation
Tayassu
Extinct genus of carnivore
species in the carnivoran family Ailuridae. It is well-represented in the Hemphillian-aged deposits at the Gray Fossil Site in Gray, Tennessee. It was significantly
Pristinailurus
Subfamily of mammals (Lutrinae)
otter (Carnivora: Mustelidae) from the late Miocene and early Pliocene (Hemphillian) of North America". Journal of Paleontology. 59 (4): 809–819. JSTOR 1304931
Otter
Genus of carnivores
years from the early Clarendonian age of the Late Miocene to the late Hemphillian age of the Early Pliocene. E. haydeni is the largest known canid of all
Epicyon
Extinct genus of ground sloths
Mylodontidae endemic to North America during the Miocene-Pliocene epochs (Hemphillian). It lived from 10.3 to 4.9 mya, existing for approximately 5.4 million
Thinobadistes
Extinct genus of ground squirrel
squirrel from North America. Fossils are known from the Blancan and Hemphillian age from localities in the United States (Nebraska, Kansas, Texas, Arizona)
Paenemarmota
First epoch of the Neogene Period
ELMA / MN (Europe) ALMA (Asia) AFLMA (Africa) (Australia) Messinian Hemphillian Montehermosan Turolian (MN 11-13) Baodean Baringian Waitean Huayquerian
Miocene
Family of mammals belonging to even-toed ungulates
Early-Late Hemphillian (HH2) faunal assemblage from Juchipila Basin, State of Zacatecas, Mexico, and its biochronologic correlation with other Hemphillian faunas
Antilocapridae
Geologic age of the late Miocene
Turolian European land mammal ages, the upper Clarendonian and lower Hemphillian North American land mammal ages and the upper Chasicoan and lower Huayquerian
Tortonian
Genus of mountain-dwelling mammal
AND ABSTRACTS, Edited by John E. Storer Shotwell, J. Arnold (1956). "Hemphillian mammalian assemblage from northeastern Oregon". Geological Society of
Pika
Family of turtles
Standish Mallory (2011). "Vertebrate paleontology of the late Miocene (Hemphillian) Wilbur Locality of central Washington". Paludicola. 8 (3): 155–185.
Emydidae
Genus of amphibians
invertebrates. The earliest known fossils of this genus are from the Hemphillian of Tennessee in the United States. Plethodon is part of the family Plethodontidae
Plethodon
Extinct species of carnivore
of canid which was endemic to North America and lived during the late Hemphillian age (between the Late Miocene and Early Pliocene). Originally described
Eucyon_ferox
Extinct genus of horse
lived in North America and Eurasia during the Late Miocene to Pliocene (Hemphillian to Blancan in the NALMA classification). They grew up to 3 feet (0.91
Cormohipparion
Extinct species of turtle
genus Sternotherus that lived in North America during the end of the Hemphillian North American land mammal age. The specific epithet of the species derives
Sternotherus_bonevalleyensis
Extinct tribe of mammals
richness during the Late Miocene and Early Pliocene. At the end of the Hemphillian (during the latest Miocene) hipparionins severely declined in diversity
Hipparionini
Extinct genus of rodents
Repomys is an extinct genus of Cricetidae that existed during the late Hemphillian and Blancan periods. Repomys arizonensis Tomida, 1987 Repomys gustelyi
Repomys
Extinct genus of bears
2005.00165.x. Dalquest, W. W. (1986). "Lower Jaw and Dentition of the Hemphillian Bear, Agriotherium (Ursidae), with the Description of a New Species"
Agriotherium
Pliocene-epoch assemblage of fossils
(Kinosternidae, Sternotherus) from the late Miocene–early Pliocene (Hemphillian) of Tennessee and Florida". Journal of Vertebrate Paleontology. 35 (1)
Gray_Fossil_Site
Geologic formation in the western United States
Group. Notably, it records the North American Land Mammal Ages (NALMAs) Hemphillian, Clarendonian, and Barstovian. It also includes an excellent record of
Ogallala_Formation
North American faunal stage according to the North American Land Mammal Ages chronology
duration of about 3.1 million years. The Blancan is preceded by the Hemphillian and followed by the Irvingtonian NALMA stages. The Blancan is named after
Blancan
Extinct genus of mammals
from modern Equus around 4.1–5.7 million years ago, during the late Hemphillian or early Blancan. Some other authors have argued that the species should
Haringtonhippus
North American faunal stage
The Clarendonian is preceded by the Barstovian and followed by the Hemphillian NALMA stages. The Clarendonian can be further divided into the substages
Clarendonian
Genus of turtles
Steve (May 2013). "Fossil Trachemys (Testudines: Emydidae) from the Late Hemphillian of Eastern Tennessee and Its Implications for the Evolution of the Emydidae"
Trachemys
Extinct genus of bears
Upper Bony Valley, Hardee and Polk counties, Florida, a typical Latest Hemphillian. The main holotype was a mandible (USNM 8838) but additional material
Huracan_(mammal)
Extinct genus of mammals
evidence suggests that it lived during the Late Miocene (Clarendonian to Hemphillian), from about 13.6 Ma to 5.3 Ma. Analysis of Protohippus' skull and teeth
Protohippus
Semiaquatic rodent
PMC 3030560. PMID 21307956. Samuels, J. X.; Zancanella, J. (2011). "An early Hemphillian occurrence of Castor (Castoridae) from the Rattlesnake Formation of Oregon"
Beaver
Extinct species of carnivore
type locality is the phosphate Palmetto Mine in Florida, which is in a Hemphillian marginal marine sandstone in the Upper Bone Valley Formation of Florida
Enhydritherium
Extinct genus of carnivores
larger animals compared to Barbourofelis. N. catocopis was found in Hemphillian rocks such as Chalk Hills Formation, Rattlesnake Formation, McKay Formation
Nimravides
Species of reptile
canebrake rattlesnake from north Florida Snakebite Parmley, Dennis (1995). "Hemphillian (Late Miocene) snakes from Nebraska, with comments on Arikareean through
Timber_rattlesnake
Extinct species of Raccoon
"A New Species of Raccoon, Procyon garberi, from Late Miocene (Late Hemphillian) Mehrten Formation, Stanislaus County, California". Bulletin of the Southern
Procyon_rexroadensis
Alkali Canyon Formation Hemphillian Chalk Butte Formation Hemphillian Drewsey Formation Hemphillian McKay Formation Hemphillian Juntura Formation Clarendonian
List of fossiliferous stratigraphic units in Oregon
List_of_fossiliferous_stratigraphic_units_in_Oregon
Extinct genus of rodents
is an extinct genus of Cricetidae that existed in Arizona during the Hemphillian period. Three species of Paronychomys are known: P. alticuspis, P. lemredfieldi
Paronychomys
Family of rodents
Mallory, V. S. (2011). "Vertebrate paleontology of the late Miocene (Hemphillian) Wilbur Locality of central Washington". Paludicola. 8 (3): 155–185.
Cricetidae
Genus of carnivores
but because of its fragmentary nature cannot be diagnosed as to species. U. webbi Tedford et al., 2009. Middle Hemphillian, Citrus County, Florida.
Urocyon
Extinct genus of mammals
Creek Formation, Nebraska Chapadmalalan Chapadmalal Formation, Argentina Hemphillian Beecher Island, Colorado Edson Beds, Kansas Rancho Viejo Beds, Mexico
Platygonus
Extinct genus of sabertooth cat
Turolian. Amphimachairodus dispersed into in North America during the Hemphillian stage, with the earliest remains being found in Withlacoochee River 4A
Amphimachairodus
Extinct genus of ground sloths
America during the Late Miocene epoch through very early Pliocene epoch (Hemphillian in the NALMA classification). Its fossils have been found in Costa Rica
Pliometanastes
Extinct species of carnivore
Borophagus littoralis Temporal range: Late Miocene (Clarendonian to Hemphillian), 11.6–9.4 Ma PreꞒ Ꞓ O S D C P T J K Pg N Scientific classification Kingdom:
Borophagus_littoralis
Extinct genus of mammals
Megatylopus Temporal range: Late Miocene-Late Pliocene (Hemphillian-Blancan) ~10.3–2.58 Ma PreꞒ Ꞓ O S D C P T J K Pg N Front limb bone of Megatylopus
Megatylopus
Subfamily of bears
Tremarctinae originate with their common ancestor, Plionarctos, in the Hemphillian faunal stage (Late Miocene) of North America, with the oldest confirmed
Tremarctinae
Sediment formation in Washington, United States
Pliocene, 4.98–4.62 million years old) River Road local fauna (late Hemphillian NALMA, latest Miocene or earliest Pliocene) Indeterminate frog or toad
Ringold_Formation
Extinct genus of ground sloths
Megalonyx (Greek, "great-claw") is an extinct genus of ground sloths of the family Megalonychidae, native to North America. It evolved during the Pliocene
Megalonyx
Geological formation in Mexico
dating back to the Neogene period. Closely corresponding to the other Hemphillian faunal assemblages of North America, it is significant for understanding
Juchipila_Formation
Extinct family of sloths
Megalonychidae Temporal range: Early Oligocene-Early Holocene (Hemphillian-Rancholabrean) (NALMA) & Tinguirirican-Lujanian (SALMA) 35–0.009 Ma PreꞒ Ꞓ
Megalonychidae
Extinct prehistoric salmon
Oncorhynchus rastrosus Temporal range: Clarendonian-Hemphillian, 12–5 Ma PreꞒ Ꞓ O S D C P T J K Pg N Skull of female specimen (UO F-55101) Reconstruction
Oncorhynchus_rastrosus
Extinct genus of mammals
Agricultural Chemical Company (Serravallian) Fort Green (Hemphillian) Payne Creek (Hemphillian) Gardinier (Zanclean) Goniodelphis was named by Allen (1941)
Goniodelphis
Extinct genus of even-toed ungulate mammals
1984). "Primitive Mylohyus (Artiodactyla: Tayassuidae) from the late Hemphillian Bone Valley of Florida". Journal of Vertebrate Paleontology. 3 (3): 152–159
Mylohyus
Extinct genus of even-toed ungulates
Synthetoceras Temporal range: Late Miocene (Hemphillian) 12.5–4.7 Ma PreꞒ Ꞓ O S D C P T J K Pg N Aqu. Burdig. Lan. Ser. Tortonian M Z P Skull of S. tricornatus
Synthetoceras
Extinct genus of mammals
relatives. A paired dental mesowear and microwear study found that early Hemphillian O. osborni had a more abrasive diet than even the merycodontines with
Osbornoceros
Extinct genus of felid
Berta, A.; Galiano, H. (1983). "Megantereon hesperus from the late Hemphillian of Florida with remarks on the phylogenetic relationships of machairodonts
Adelphailurus
back to the Serravallian to earliest Pliocene stages (Clarendonian, Hemphillian and earliest Blancan in the NALMA classification) of the Neogene period
Goliad_Formation
Geologic timescale for prehistoric Asian fauna
Blancan Jinglean Early Pliocene Late Hemphillian–early Blancan Baodean Late Miocene Late Clarendonian–early Hemphillian Bahean ~11.1 Ma Middle–late Miocene
Asian_land_mammal_age
Geologic formation in Oregon, United States
The Rattlesnake Formation is a Miocene to late Pliocene geologic formation found along the John Day River Valley of Oregon, in the Western United States
Rattlesnake_Formation
Species of canid
42. The gray fox appeared in North America during the mid-Pliocene (Hemphillian land animal age) epoch 3.6 million years ago (AEO) with the first fossil
Gray_fox
Lake in the U.S. state of Nevada
Göhlich (2021): New mammutids (Proboscidea) from the Clarendonian and Hemphillian of Oregon – a survey of Mio-Pliocene mammutids from North America v t
Massacre_Lake
Geological formation in Florida
Neogene Epoch: Miocene to Pliocene Faunal stage: Arikareean through Hemphillian, ~23.03–4.9 mya, calculates to a period of ~18.13 million years The Peace
Peace River Formation (Florida)
Peace_River_Formation_(Florida)
Genus of turtle
1903 - Pliocene †Clemmys owyheensis Brattstrom & Sturn, 1959 - Miocene (Hemphillian) Feldman, C.R. and J.F. Parham. (2002). Molecular phylogenetics of Emydine
Clemmys
Extinct species of turtle
(Kinosternidae, Sternotherus ) from the late Miocene–early Pliocene (Hemphillian) of Tennessee and Florida". Journal of Vertebrate Paleontology. 35 (1)
Sternotherus_palaeodorus
Geologic formation in coastal southern Orange County, California
the Niguel Formation. The Oso Member has been dated to the early late Hemphillian (6.6 to 5.8 Ma) based on the presence of Dinohippus interpolatus and
Capistrano_Formation
(1984). "Astrohippus and Dinohippus from the Yepómera local fauna (Hemphillian, Mexico) and implications for the phylogeny of one-toed horses". Journal
Evolution_of_the_horse
Extinct genus of mammal
Donald D. De Blieux (2010). "Rodents and Lagomorphs (Mammalia) from the Hemphillian (Late Miocene) of Utah". Journal of Vertebrate Paleontology. 30 (1):
Metaliomys
Extinct genus of mammal
Science. 15: 120–150. Carranza-Castañeda, O.; Miller, W. E. (May 1996). "Hemphillian and Blancan Felids from central Mexico". Journal of Paleontology. 70
Miracinonyx
Zoo in Asheville, North Carolina
"A new species of Teleoceras (Mammalia, Rhinocerotidae) from the Late Hemphillian of Tennessee" (PDF). Bulletin of the Florida Museum of Natural History
Western North Carolina Nature Center
Western_North_Carolina_Nature_Center
Geologic formation in the southern United States
The Citronelle Formation is a Hemphillian geologic formation in Alabama, Florida, Mississippi and Louisiana. It preserves fossils of: Nannippus cf. lenticularis
Citronelle_Formation
turtle (Testudines: Emydidae: Deirochelyinae: Trachemys) from the late Hemphillian (late Miocene/early Pliocene) of eastern Tennessee and the evolution
Trachemys_haugrudi
Geologic formation in Wyoming, United States
refined its placement to the late Miocene (late Barstovian to early Hemphillian ages). U-Pb zircon dating of interbedded volcanic ashes within the formation
Moonstone_Formation
Extinct genus of mammals
shorelines. They thrived approximately 5 million years ago during the Hemphillian age. A map of locations where fossils of Nanosiren garciae has been created
Nanosiren
Extinct genus of deer
Cranioceras Temporal range: Barstovian–Hemphillian ~15.97–5.3 Ma PreꞒ Ꞓ O S D C P T J K Pg N Life restoration of Cranioceras unicornis Scientific classification
Cranioceras
Genus of turtle
†Hesperotestudo gilbertii (Hay, 1899) Ogallala Formation, Kansas, Hemphillian, Late Miocene-Early Pliocene (Tortonian–Zanclean) †Hesperotestudo orthopygia
Hesperotestudo
Geologic formation in Panama
formation dates to the latest Serravallian to Tortonian (Clarendonian to Hemphillian in the NALMA classification), from 12 to 8.5 Ma. It preserves many fossils
Gatún_Formation
Geologic formation in the Mojave Desert, California, U.S.
P. Whistler (2014). "New late Miocene (latest Clarendonian to early Hemphillian) cricetid rodents from the upper part of the Dove Spring Formation, Mojave
Dove_Spring_Formation
Group of extinct animals
replacing Machairodus. It would also disperse into North America during the Hemphillian stage, with the oldest fossils being found in Withlacoochee River 4A
Saber-toothed_predator
Subspecies of mammal
they form the tribe Antilocaprini. Texoceros first occurs in the late Hemphillian and disappears by the Blancan age. The modern pronghorn first appears
Mexican_pronghorn
Geologic formation in New Mexico, U.S.
chemically resembles. This helps establish the absolute date for the late Hemphillian, since a fossil assemblage of that stage is found in the zone. The Lower
Chamita_Formation
Genus of reptile
turtle (Testudines: Emydidae: Deirochelyinae: Trachemys) from the late Hemphillian (late Miocene/early Pliocene) of eastern Tennessee and the evolution
Chrysemys
Geologic formation in Panama
the Middle to Late Miocene (Tortonian to Messinian, Clarendonian to Hemphillian in the NALMA classification) period. The Chagres Formation is exposed
Chagres_Formation
Extinct genus of mammals
Florida Museum of Natural History. MacFadden, Bruce J. (1986). "Late Hemphillian Monodactyl Horses (Mammalia, Equidae) from the Bone Valley Formation
Neohipparion
Extinct subfamily of birds
Mancalla was a common species throughout the Pliocene, appearing in the Hemphillian stage of the Late Miocene (5–9 mya), and spreading in the Pliocene, with
Mancallinae
Genus of turtles
turtle (Testudines: Emydidae: Deirochelyinae: Trachemys) from the late Hemphillian (late Miocene/early Pliocene) of eastern Tennessee and the evolution
Deirochelys
Extinct genus of skunk
mustelid from a fragmentary lower jaw found in the Edson Quarry, from late Hemphillian Kansas. It was described as the type of the new genus. Due to its fragmentary
Martinogale
Extinct genus of mammals
Protolabis Temporal range: Arikareean-Hemphillian ~20.4–5.3 Ma PreꞒ Ꞓ O S D C P T J K Pg N Protolabis gracilis Scientific classification Kingdom: Animalia
Protolabis
Extinct genus of carnivores
Bennet, D. K. (1979). "The fossil fauna from Lost and Found Quarries (Hemphillian: latest Miocene), Wallace County, Kansas". Occasional Papers of the Museum
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